Scarce Swallowtail
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The scarce swallowtail (''Iphiclides podalirius'') is a butterfly belonging to the family
Papilionidae Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the larges ...
. It is also called the sail swallowtail or pear-tree swallowtail.


Subspecies

Subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
include:"''Iphiclides'' Hübner, [1819]"
at Markku Savela's ''Lepidoptera and Some Other Life Forms''
*''Iphiclides podalirius podalirius'' (Central and Southern Europe) *''Iphiclides podalirius feisthamelii'' (North Africa, Spain and southwest France) *''Iphiclides podalirius persica'' Verity, 1911 *''Iphiclides podalirius virgatus'' (Butler, 1865) ''Iphiclides podalirius feisthamelii'' is sometimes treated as a valid species, ''Iphiclides feisthamelii''.


Distribution

Despite the name "scarce swallowtail", this species is quite common. The scarcity of United Kingdom migrants is responsible for the English vernacular name. This species is widespread in the East
Palearctic realm The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
and in most of Europe with the exception of the northern parts. Its range extends northwards to
Lower Lusatia Lower Lusatia (; ; ; szl, Dolnŏ Łużyca; ; ) is a historical region in Central Europe, stretching from the southeast of the German state of Brandenburg to the southwest of Lubusz Voivodeship in Poland. Like adjacent Upper Lusatia in the sou ...
and central Poland and eastwards across
Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
and
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...
as far as the
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
, Pakistan, India and western China. A few specimens of the scarce swallowtail have been reported from central Sweden and the United Kingdom but they were probably only strays and not migrants.


Habitat

These
swallowtail butterflies Swallowtail butterflies are large, colorful Butterfly, butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes ...
inhabit
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate both ...
s, towns as well as the countryside, in
fields Fields may refer to: Music *Fields (band), an indie rock band formed in 2006 *Fields (progressive rock band), a progressive rock band formed in 1971 * ''Fields'' (album), an LP by Swedish-based indie rock band Junip (2010) * "Fields", a song by ...
and open
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s. They are found in places with sloe thickets and particularly
orchard An orchard is an intentional plantation of trees or shrubs that is maintained for food production. Orchards comprise fruit- or nut-producing trees which are generally grown for commercial production. Orchards are also sometimes a feature of larg ...
s. In the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
they can be found up to altitudes of 2000 m, but usually they prefer foothills and lower levels.Euro Butterflies by Matt Rowlings
/ref> The presence of ''Iphiclides podalirius'' in the floodplain of the
Morava River Morava may refer to: Rivers * Great Morava (''Velika Morava''; or only Morava), a river in central Serbia, and its tributaries: ** South Morava (''Južna Morava'') *** Binač Morava (''Binačka Morava'') ** West Morava (''Zapadna Morava'') * Mo ...
in the
Slovak Republic Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
have been found to be a good indicator of relatively well preserved xerothermic
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
habitats with forest-steppe vegetation, which have no cutting history.


Status

In some years the scarce swallowtail is quite abundant. The scarce swallowtail is getting rarer as blackthorn bushes are being cleared. The butterfly is now protected by law in the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
,
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
,
Luxembourg Luxembourg ( ; lb, Lëtzebuerg ; french: link=no, Luxembourg; german: link=no, Luxemburg), officially the Grand Duchy of Luxembourg, ; french: link=no, Grand-Duché de Luxembourg ; german: link=no, Großherzogtum Luxemburg is a small lan ...
,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
,
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
and
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
. It is considered rare and endangered and protected in some provinces of Austria and of indeterminate status throughout Europe. In Armenia the species demonstrates stable population trend and is evaluated as Least Concern. Though referred by some authorities to be of status "vulnerable", it is however unlisted in the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
.


Description

''Iphiclides podalirius'' has a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
of in males, of in females. It is a very large distinctive butterfly. The background color of the wings is creamy white or pale yellow. On the front wings there are six tiger stripes and wedge-shaped markings. At the outer edge of the hind wings there are blue crescent markings, with an oblong, orange spot at the back corner and a relatively long tail. This species is rather similar to ''
Papilio machaon ''Papilio machaon'', the Old World swallowtail, is a butterfly of the family Papilionidae. The butterfly is also known as the common yellow swallowtail or simply the swallowtail (a common name applied to all members of the family, but this speci ...
'', '' Papilio hospiton'', ''
Papilio alexanor ''Papilio alexanor'', the Alexanor or southern swallowtail, is a butterfly species in the Papilionidae or swallowtails. The insect has a wingspan of 62–70 mm. It flies from April to July in a single generation, in mountainous regions (170 ...
'' and ''
Protographium marcellus ''Eurytides marcellus'', the zebra swallowtail, (formerly listed under genera ''Protographium'', ''Iphiclides'', '' Graphium'' and ''Papilio'' by some authorities) is a swallowtail butterfly native to the eastern United States and south-eastern ...
''. Iphiclides podalirius MHNT CUT 2013 3 9 Male Dos Cahors.jpg, Male Iphiclides podalirius MHNT CUT 2013 3 9 Male Ventre Cahors.jpg, Male underside Iphiclides podalirius MHNT CUT 2013 3 9 Female Dos Cahors.jpg, Female Iphiclides podalirius MHNT CUT 2013 3 9 Female Ventre Cahors.jpg, Female underside


Biology

Adults of ''Iphiclides podalirius'' fly from March to October. There are one, two, or three generations a year. Caterpillars are
polyphagous Feeding is the process by which organisms, typically animals, obtain food. Terminology often uses either the suffixes -vore, -vory, or -vorous from Latin ''vorare'', meaning "to devour", or -phage, -phagy, or -phagous from Greek φαγε ...
feeding on ''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' (Finland), ''
Crataegus monogyna ''Crataegus monogyna'', known as common hawthorn, one-seed hawthorn, or single-seeded hawthorn, is a species of flowering plant in the rose family Rosaceae. It is native to Europe, northwestern Africa, and West Asia, but has been introduced in ...
'' (Palaearctic), ''
Malus pumila An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus ''Malus''. The tree originated in Central Asia, where its wild ancestor, ' ...
'' and ''
Malus domestica An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus ''Malus''. The tree originated in Central Asia, where its wild ancestor, ' ...
'' (Palaearctic), ''Prunus'' (Finland, Palaearctic), ''
Prunus armeniaca ''Prunus armeniaca'' is the most commonly cultivated apricot species. The native range is somewhat uncertain due to its extensive prehistoric cultivation. Genetic studies indicate Central Asia is the center of origin. It is extensively cultivated ...
'' (Palaearctic, Spain), ''
Prunus avium ''Prunus avium'', commonly called wild cherry, sweet cherry, gean, or bird cherryWorld Economic Plants: A Standard Reference, Second Edition'. CRC Press; 19 April 2016. . p. 833–. is a species of cherry, a flowering plant in the rose family, ...
'' (Palaearctic, Spain) ''
Prunus cerasus ''Prunus cerasus'' (sour cherry, tart cherry, or dwarf cherry) is a species of ''Prunus'' in the subgenus '' Cerasus'' ( cherries), native to much of Europe and southwest Asia. It is closely related to the sweet cherry (''Prunus avium''), but ...
'' (Palaearctic, Russia), ''
Prunus domestica ''Prunus domestica'', the European plum is a species of flowering plant in the family Rosaceae. A deciduous tree, it includes many varieties of the fruit trees known as plums in English, though not all plums belong to this species. The greengag ...
'' (Palaearctic), ''
Prunus dulcis The almond (''Prunus amygdalus'', syn. ''Prunus dulcis'') is a species of tree native to Iran and surrounding countries, including the Levant. The almond is also the name of the edible and widely cultivated seed of this tree. Within the genus ...
'' (Palaearctic, Spain), ''
Prunus padus ''Prunus padus'', known as bird cherry, hackberry, hagberry, or Mayday tree, is a flowering plant in the rose family. It is a species of cherry, a deciduous small tree or large shrub up to tall. It is the type species of the subgenus '' Padus' ...
'' (Palaearctic), ''
Prunus persica ''Prunus'' is a genus of trees and shrubs, which includes (among many others) the fruits plums, cherries, peaches, nectarines, apricots, and almonds. Native to the North American temperate regions, the neotropics of South America, and the paleo ...
'' (Palaearctic), ''
Prunus spinosa ''Prunus spinosa'', called blackthorn or sloe, is a species of flowering plant in the rose family Rosaceae. The species is native to Europe, western Asia, and regionally in northwest Africa. It is locally naturalized in New Zealand, Tasmania, ...
'' (Palaearctic), ''
Pyrus Pears are fruits produced and consumed around the world, growing on a tree and harvested in the Northern Hemisphere in late summer into October. The pear tree and shrub are a species of genus ''Pyrus'' , in the family Rosaceae, bearing the po ...
'' (Palaearctic) and ''
Sorbus aucuparia ''Sorbus aucuparia'', commonly called rowan (UK: /ˈrəʊən/, US: /ˈroʊən/) and mountain-ash, is a species of deciduous tree or shrub in the rose family. It is a highly variable species, and botanists have used different Circumscription (taxo ...
'' (Palaearctic). The caterpillars spin little pads on leaves and grip them firmly. The newly hatched caterpillar is dark in colour with two smaller and two bigger greenish patches on the dorsal side; later they are greenish with yellowish dorsal and side stripes. The summer chrysalids are green as a rule, the hibernating ones are brown. A number of hibernating chrysalids fall prey to various enemies. The caterpillars of the scarce swallowtail have been noted to leave silk trails from the permanent resting sites to feeding sites. This has been seen in both solitary and territorial larvae with larvae having the ability to discern their trails from those of others. Research on pupae of ''Iphiclides podalirius'' in Spain indicates that the pupae manifest in two colours, green and brown, for the purpose of camouflage. The green pupae develop on host plants and develop directly while brown pupae enter into
diapause In animal dormancy, diapause is the delay in development in response to regular and recurring periods of adverse environmental conditions.Tauber, M.J., Tauber, C.A., Masaki, S. (1986) ''Seasonal Adaptations of Insects''. Oxford University Press It ...
in the leaf litter. Pupating larvae tend to form green pupae before August while after August they tend to form brown pupae. Duration of the photophase or light period appears to be the mechanism which dictates the path of development of the pupa. The results suggest that the green pupa develop on food plants to avoid predation by small mammals and visual avian predators while the brown pupa develop on leaf litter to avoid avian predators.


Life cycle

File: Iphiclides podalirius 1.jpg, Caterpillar File:Iphiclides podalirius pupating.jpg, Pupating caterpillar File:SailSwallowtail.jpg, Imago


References


External links


Scarce swallowtail
on Butterflycorner.net
Lepiforum.de
* Paolo Mazzei, Daniel Morel, Raniero Panfil

* ttps://www.guypadfield.com/scarceswallowtail.html ''Iphiclides podalirius'' on Guy Padfield's Butterfly Page* Butterfly Conservation Armeni

{{Taxonbar, from=Q31080 Iphiclides Butterflies of Asia Butterflies of Europe Butterflies described in 1758 Taxa named by Carl Linnaeus