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''Scaevola taccada'', also known as beach cabbage, sea lettuce, or beach naupaka, is a
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the family
Goodeniaceae Goodeniaceae is a family of flowering plants in the order Asterales. It contains about 404 species in twelve genera. The family is distributed mostly in Australia, except for the genus '' Scaevola'', which is pantropical. Its species are found ac ...
found in coastal locations in the tropical areas of the Indo-Pacific. It is a common beach shrub throughout the
Arabian Sea The Arabian Sea ( ar, اَلْبَحرْ ٱلْعَرَبِيُّ, Al-Bahr al-ˁArabī) is a region of the northern Indian Ocean bounded on the north by Pakistan, Iran and the Gulf of Oman, on the west by the Gulf of Aden, Guardafui Channel ...
, the tropical
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by t ...
and the tropical islands of the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contin ...
. There has long been confusion over the correct scientific name for this species. It is also known by the synonym ''Scaevola sericea''.


Description

''Scaevola taccada'' is a large bush reaching up to about 4 m in height typical of
littoral zone The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal a ...
s where it grows very close to the sea exposed to the salt spray, usually on sandy or pebbly soils. Leaves are slightly succulent, about 20 cm long, closely alternate and crowded at the stem tips. They are
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
with a fleshy-looking yellowish green color. The fruits and flowers are white. ''Scaevola taccada'' blooms the whole year round and the flowers have a fan-like shape which gives them the name fanflower or half flower. The fruits float in seawater and are propagated by
ocean current An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth contours ...
s, this bush being one of the
pioneer plant Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire. Pioneer flora Some lichens grow on rocks without soil, so m ...
s in new sandbanks in tropical areas. ''Scaevola taccada'' is similar to ''
Scaevola plumieri ''Scaevola plumieri'' (common name gullfeed) is a species of plant in the family Goodeniaceae which grows on coastal dunes in the tropics and subtropics. Description ''Scaevola plumieri'' is a many branched evergreen shrub, which has succulent h ...
'' – both were included in the original
circumscription Circumscription may refer to: *Circumscribed circle * Circumscription (logic) *Circumscription (taxonomy) *Circumscription theory, a theory about the origins of the political state in the history of human evolution proposed by the American anthrop ...
of the species. ''S. taccada'' has evident calyx lobes and white fruit on which the calyx lobes persist. ''S. plumieri'' has short or absent lobes on its calyx and black fruit when ripe. File:Scaevola taccada fruit.jpg, White fruit of ''S. taccada'' with persistent calyx lobes File:Scaevola plumieri 2.jpg, Black fruit of ''S. plumieri''


Taxonomy

Two shrubby ''Scaevola'' species occur along the coasts of tropical and subtropical regions of the world. They sometimes occur in the same region, but one is more western, reaching Atlantic coasts, and one more eastern, reaching out into the Pacific.
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...
initially included both species in his ''Lobelia plumieri'', which he later considered to be the sole species in his new genus ''Scaevola'' (although the combination ''Scaevola plumieri'' was first published by Martin Vahl). There has been confusion for many years over the correct name of the two species when they are recognized as distinct. ''
Scaevola plumieri ''Scaevola plumieri'' (common name gullfeed) is a species of plant in the family Goodeniaceae which grows on coastal dunes in the tropics and subtropics. Description ''Scaevola plumieri'' is a many branched evergreen shrub, which has succulent h ...
'' (L.) Vahl is now used as the name for the western species. The earliest name now recognized as applying to the eastern species, ''Lobelia taccada'', was published by
Joseph Gaertner Joseph Gaertner (12 March 1732 – 14 July 1791) was a German botanist, best known for his work on seeds, ''De Fructibus et Seminibus Plantarum'' (1788-1792). Biography He was born in Calw, and studied in Göttingen under Albrecht von Halle ...
in 1788.
William Roxburgh William Roxburgh FRSE FRCPE FLS (3/29 June 1751 – 18 February 1815) was a Scottish surgeon and botanist who worked extensively in India, describing species and working on economic botany. He is known as the founding father of Indian botany. ...
indirectly referred to this name when transferring it to the genus '' Scaevola'' in 1788. Separately, and later, Martin Vahl described ''Scaevola sericea'' in 1791, based on a specimen from
Niue Niue (, ; niu, Niuē) is an island country in the South Pacific Ocean, northeast of New Zealand. Niue's land area is about and its population, predominantly Polynesian, was about 1,600 in 2016. Niue is located in a triangle between Tong ...
, a small island in the south Pacific Ocean. In 1980, Jeffrey argued that the correct name for the species was ''Scaevola sericea'', since Roxburgh's transfer was not acceptable under the nomenclature code. However,
Green Green is the color between cyan and yellow on the visible spectrum. It is evoked by light which has a dominant wavelength of roughly 495570 nm. In subtractive color systems, used in painting and color printing, it is created by a combi ...
in 1991 considered that Jeffrey was mistaken, the transfer being valid, so that the correct name for the eastern species was ''Scaevola taccada'', the name used, for example, by the online ''Flora of China''. The '' International Plant Names Index'' accepts this analysis.


Distribution

The species is found in coastal areas and beaches of
Okinawa is a prefecture of Japan. Okinawa Prefecture is the southernmost and westernmost prefecture of Japan, has a population of 1,457,162 (as of 2 February 2020) and a geographic area of 2,281 km2 (880 sq mi). Naha is the capital and largest city ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
,
Southern China South China () is a geographical and cultural region that covers the southernmost part of China. Its precise meaning varies with context. A notable feature of South China in comparison to the rest of China is that most of its citizens are not n ...
,
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
,
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
,
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
,
East Timor East Timor (), also known as Timor-Leste (), officially the Democratic Republic of Timor-Leste, is an island country in Southeast Asia. It comprises the eastern half of the island of Timor, the exclave of Oecusse on the island's north-west ...
,
Northern Australia The unofficial geographic term Northern Australia includes those parts of Queensland and Western Australia north of latitude 26° and all of the Northern Territory. Those local government areas of Western Australia and Queensland that lie p ...
,
Polynesia Polynesia () "many" and νῆσος () "island"), to, Polinisia; mi, Porinihia; haw, Polenekia; fj, Polinisia; sm, Polenisia; rar, Porinetia; ty, Pōrīnetia; tvl, Polenisia; tkl, Polenihia (, ) is a subregion of Oceania, made up of ...
,
Melanesia Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from Indonesia's New Guinea in the west to Fiji in the east, and includes the Arafura Sea. The region includes the four independent countries of Fiji, Va ...
,
Micronesia Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, and ...
, East Africa,
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
,
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label= Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It ...
,
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, ...
,
Oman Oman ( ; ar, عُمَان ' ), officially the Sultanate of Oman ( ar, سلْطنةُ عُمان ), is an Arabian country located in southwestern Asia. It is situated on the southeastern coast of the Arabian Peninsula, and spans the mouth of ...
,
Yemen Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in Western Asia. It is situated on the southern end of the Arabian Peninsula, and borders Saudi Arabia to the Saudi Arabia–Yemen border, north and ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Maldives Maldives (, ; dv, ދިވެހިރާއްޖެ, translit=Dhivehi Raajje, ), officially the Republic of Maldives ( dv, ދިވެހިރާއްޖޭގެ ޖުމްހޫރިއްޔާ, translit=Dhivehi Raajjeyge Jumhooriyyaa, label=none, ), is an archipelag ...
,
Burma Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
,
Chagos Islands The Chagos Archipelago () or Chagos Islands (formerly the Bassas de Chagas, and later the Oil Islands) is a group of seven atolls comprising more than 60 islands in the Indian Ocean about 500 kilometres (310 mi) south of the Maldives arc ...
, Comoros, and Réunion. In the United States (Florida, Puerto Rico, US Virgin Islands) as well as many other Caribbean nations and the Bahamas, ''Scaevola taccada'' has become an invasive species, pushing away the native Caribbean ''
Scaevola plumieri ''Scaevola plumieri'' (common name gullfeed) is a species of plant in the family Goodeniaceae which grows on coastal dunes in the tropics and subtropics. Description ''Scaevola plumieri'' is a many branched evergreen shrub, which has succulent h ...
'' species from its native habitat.


Habitat

''Scaevola taccada'' typically grows directly on the beaches of tropical coasts with a preference for beach crests on coral sands. It grows within the salt spray area and it is amongst the first pioneer plant colonisers on tropical atolls and sandbanks. Besides seeds, it is easily propagated from cuttings. It prefers well drained sandy soils and it is a very salt tolerant scrub. ''Scaevola taccada'' is sometimes found growing in loose plant communities with
coconut palm The coconut tree (''Cocos nucifera'') is a member of the palm tree family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or t ...
s, soldierbush, beach morning glory, beach gardenia, several pandanus species, beach calophyllum followed by
portia tree ''Thespesia populnea'', commonly known as the portia tree (), Pacific rosewood, Indian tulip tree, or milo, among other names, is a species of flowering plant belonging to the mallow family, Malvaceae. It is a tree found commonly on coasts ar ...
, sea almond, beach hibiscus, '' Cordia subcordata'' and others. The plant is often featured prominently on tropical island postcards and wallpapers.


Use

In some islands of the Pacific, ''Scaevola taccada'' is used to prevent
coastal erosion Coastal erosion is the loss or displacement of land, or the long-term removal of sediment and rocks along the coastline due to the action of waves, currents, tides, wind-driven water, waterborne ice, or other impacts of storms. The landwa ...
as well as for landscaping. It is also planted on the beach crests to protect other cultivated plants from the salt spray. Parts of the plant are also used in
Polynesia Polynesia () "many" and νῆσος () "island"), to, Polinisia; mi, Porinihia; haw, Polenekia; fj, Polinisia; sm, Polenisia; rar, Porinetia; ty, Pōrīnetia; tvl, Polenisia; tkl, Polenihia (, ) is a subregion of Oceania, made up of ...
n and Asian
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
. Drops from the plant were used in cases of eye irritation by Chamorro breath-hold spearfishermen in the Mariana Islands. Historically in the
Maldives Maldives (, ; dv, ދިވެހިރާއްޖެ, translit=Dhivehi Raajje, ), officially the Republic of Maldives ( dv, ދިވެހިރާއްޖޭގެ ޖުމްހޫރިއްޔާ, translit=Dhivehi Raajjeyge Jumhooriyyaa, label=none, ), is an archipelag ...
the leaves of this plant were often used as famine food.Eating on the Islands - As times have changed, so has the Maldives' unique cuisine and culture
/ref>


Gallery


References


External links


''Scaevola sericea'' occurrence data
from GBIF
Merambong or Sea lettuce - ''Scaevola taccada''
* * {{Taxonbar, from1=Q2702446, from2=Q73291738, from3=Q50845588 taccada Flora of Hawaii Flora of Queensland Asterales of Australia Flora of tropical Asia Flora of Oman Flora of Tanzania Leaf vegetables