Saloumia
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''Saloumia'' is an extinct genus of the order
Proboscidea The Proboscidea (; , ) are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From ...
. It is one of the oldest members of the order and lived in the middle
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ðž ...
. It is known only from a single molar, whose pronounced bumpy chewing surface indicates it is probably closely related to ''
Moeritherium ''Moeritherium'' ("the beast from Lake Moeris") is an extinct genus of primitive proboscideans. These prehistoric mammals are related to the elephant and, more distantly, sea cows and hyraxes Hyraxes (), also called dassies, are small, thick ...
''.


Description

The genus is known only from a partial upper molar 10.7 mm long and 12.3 mm wide, missing the posterior section. The molar is smaller than those of ''Moeritherium'', but larger than those of ''
Eritherium ''Eritherium'' is an extinct genus of early Proboscidea found in the Ouled Abdoun basin (early Thanetian age), Morocco. It lived about 60 million years ago. It was first named by Emmanuel Gheerbrant in 2009 and the type species is ''Eritherium ...
'' and ''
Phosphatherium ''Phosphatherium escuillei'' is a basal proboscidean that lived from the Late Paleocene to the early stages of the Ypresian age until the early Thanetian some 56 million years ago in North Africa. Research has suggested that ''Phosphatherium'' ex ...
''. The tooth was low-crowned, more clearly than that of ''Moeritherium''. The masticatory surface had four cusps. However, their tips were apparently broken off before fossilization. The four cusps each formed two pairs, which were arranged perpendicular to the longitudinal axis of the tooth, giving it a
bilophodont The molars or molar teeth are large, flat tooth, teeth at the back of the mouth. They are more developed in mammal, mammals. They are used primarily to comminution, grind food during mastication, chewing. The name ''molar'' derives from Latin, ...
structure characteristic of early proboscideans. The front pair of cusps (the paracone and the protocone) had no additional ridges. The bases of the two cusps were connected to each other, and a side cusp formed a paraconule. The rear pair of cusps (the metacone and hypocone) are damaged, but the hypocone was originally relatively central and was very large, its dimensions similar to those of the protocone. Its inward shifted position differs from the more marginal position of ''Moeritheriums hypocone. There was no metaconule on the second pair of cusps. The central longitudinal groove, which divides the tooth into two halves was only weakly developed. A well-developed cingulum was present on the tongue side. The presence and position of the cingulum distinguishes ''Saloumia'' from ''Moeritherium''. On both the tongue and cheek sides, the cingulum merged into a shearing edge. The enamel as a whole was thick and heavily grooved.


Discovery and naming

The sole specimen of ''Saloumia'' was found in the Lam-Lam Formation near the village of M’Bodione Dadere, north of the city of
Kaolack Kaolack ( ar, كاولاك; wo, Kawlax) is a town of 172,305 people (2002 census) on the north bank of the Saloum River and the N1 road in Senegal. It is the capital of the Kaolack Region, which borders The Gambia to the south. Kaolack is an i ...
in the central-western part of Senegal. The landscape in the region is relatively flat and is crossed by the
Saloum River The Saloum River rises about 105 kilometers east of Kaolack, Senegal, and flows into the Atlantic Ocean. The significant Saloum Delta is located at its mouth, which is protected as Saloum Delta National Park. The river basin lies within the Serer ...
. The area contains several meters of Quaternary sands. Access to fossil outcrops is usually by means of well shafts and boreholes. The Lam-Lam formation consists of a limestone rich in mollusks and sea urchins, which alternates with a clayey limestone and marl. The limestones contain various
foraminiferans Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an ...
, of which '' Globigerinatheca'', '' Cassigerinelloita'' and '' Pseudohastigerina'' are typical of the Lutetian, around 44 million years ago. Other vertebrates from the Lam-Lam formation are mostly ray-finned fishes. The overlying Taïba Formation contained the remains of an
archaeocete Archaeoceti ("ancient whales"), or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene (). Representing the earliest cetacean Evolutionary radiation, radiation, t ...
whale. The generic name is derived from the Saloum River. The specific epithet honors the geologist
Alexandre Gorodiski Alexandre may refer to: * Alexandre (given name) * Alexandre (surname) * Alexandre (film) See also * Alexander Alexander is a male given name. The most prominent bearer of the name is Alexander the Great, the king of the Ancient Greek kingdom o ...
, who discovered the tooth in 1952 during mapping work. Gorodiski published the find with René Lavocat a year later, describing it as a species of ''Moeritherium''.


Further reading

* Emmanuel Gheerbrant, Baadi Bouya and Mbarek Amaghzaz: ''Dental and cranial anatomy of Eritherium azzouzorum from the Paleocene of Morocco, earliest known proboscidean material.'' Palaeontographica, Abteilung A 297 (5/6), 2012, S. 151–183. *Jeheskel Shoshani and Pascal Tassy: ''Advances in proboscidean taxonomy & classification, anatomy & physiology, and ecology & behavior.'' Quaternary International 126–128, 2005, S. 5–20 *Emmanuel Gheerbrant: ''Paleocene emergence of elephant relatives and the rapid radiation of African ungulates.'' PNAS 106 (6), 2009, S. 10717–10721. *Jeheskel Shoshani, W. J. Sanders and Pascal Tassy: ''Elephants and other Proboscideans: a summary of recent findings and new taxonomic suggestions.'' In: G. Cavarretta et al. (Hrsg.): ''The World of Elephants – International Congress. Consiglio Nazionale delle Ricerche.'' Rom, 2001, S. 676–679


References

Rodolphe Tabuce, Raphaël Sarr, Sylvain Adnet, Renaud Lebrun, Fabrice Lihoreau, Jeremy E. Martin, Bernard Sambou, Mustapha Thiam und Lionel Hautier: ''Filling a gap in the proboscidean fossil record: a new genus from the Lutetian of Senegal.'' Journal of Paleontology, 2019, doi:10.1017/jpa.2019.98 Maureen A. O'Leary, Raphaël Sarr, Raymond Malou, El Hdji Sow, Christopher J. Lepre und Robert V. Hill: ''A New Fossil Amiid from the Eocene of Senegal and the Persistence of Extinct Marine Amiids after the Cretaceous–Paleogene Boundary.'' Copeia 4, 2012, S. 603–608 Phillip D. Gingerich: ''Cetacea.'' In: Lars Werdelin und William Joseph Sanders (Hrsg.): ''Cenozoic Mammals of Africa.'' University of California Press, Berkeley, London, New York, 2010, S. 873–899 {{Taxonbar, from=Q97380035 Eocene proboscideans Prehistoric placental genera