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The green swordtail (''Xiphophorus hellerii'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
/
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
in
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Poeciliidae The Poeciliidae are a family of freshwater fishes of the order Cyprinodontiformes, the tooth-carps, and include well-known live-bearing aquarium fish, such as the guppy, molly, platy, and swordtail. The original distribution of the family was t ...
of
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Cyprinodontiformes. A live-bearer, it is closely related to the southern platyfish or 'platy' (''X. maculatus'') and can crossbreed with it. It is native to an area of
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...
and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
stretching from
Veracruz Veracruz (), formally Veracruz de Ignacio de la Llave (), officially the Free and Sovereign State of Veracruz de Ignacio de la Llave ( es, Estado Libre y Soberano de Veracruz de Ignacio de la Llave), is one of the 31 states which, along with Me ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, to northwestern
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
.


Description

The male green swordtail grows to a maximum overall length of and the female to . The name 'swordtail' is derived from the elongated lower lobe of the male's
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
(tailfin).
Sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
is moderate, with the female being larger than the male, but lacking the 'sword'. The wild form is olive green in color, with a red or brown lateral stripe and speckles on the dorsal and, sometimes, caudal fins. The male's 'sword' is yellow, edged in black below. Captive breeding has produced many color varieties, including black, red, and many patterns thereof, for the aquarium hobby. All varieties share the dark-red or brown central stripe. The green swordtail prefers swift-flowing, heavily vegetated rivers and streams, but is also found in warm springs and canals. Omnivorous, its diet includes both plants and small
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s,
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s, and
annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecol ...
worms.


Invasive species

''X. hellerii'' has become a nuisance pest as an
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
in a number of countries. It has caused ecological damage because of its ability to rapidly reproduce in high numbers. Feral populations have established themselves in southern
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, including Natal,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
,
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
and eastern
Transvaal Transvaal is a historical geographic term associated with land north of (''i.e.'', beyond) the Vaal River in South Africa. A number of states and administrative divisions have carried the name Transvaal. * South African Republic (1856–1902; af, ...
in
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
and
Otjikoto Lake Otjikoto Lake is the smaller of only two permanent natural lakes in Namibia. It is a sinkhole lake that was created by a collapsing Karst topography, karst cave. It is located north-west of Tsumeb and only 100 meters from the main road B1 road (Na ...
in
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
. Significant populations have also established themselves along the east coast of
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
.


Etymology and taxonomy

The
binomial Binomial may refer to: In mathematics *Binomial (polynomial), a polynomial with two terms * Binomial coefficient, numbers appearing in the expansions of powers of binomials *Binomial QMF, a perfect-reconstruction orthogonal wavelet decomposition ...
of the green swordtail is made up of its generic name, ''Xiphophorus'' which means ''sword bearer'' in Greek, a reference to the extended lower part of the caudal fin which is referred to as the "sword". The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honours the collector of the type, the
Austrian Austrian may refer to: * Austrians, someone from Austria or of Austrian descent ** Someone who is considered an Austrian citizen, see Austrian nationality law * Austrian German dialect * Something associated with the country Austria, for example: ...
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
Karl Bartholomaeus Heller (1824–1880), who discovered this fish while exploring México in 1845–1848. The green swordtail was described from Heller's type by Johann Jakob Heckel in 1848 with the type locality given as Orizaba, Mexico. It is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of the genus ''
Xiphophorus ''Xiphophorus'' is a genus of euryhaline and freshwater fishes in the family Poeciliidae of order Cyprinodontiformes, native to Mexico and northern Central America. The many ''Xiphophorus'' species are all known as platyfish (or platies) and s ...
''.


Aquarium fish

One of the most popular
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
fish, the green swordtail has been bred into various hybrid forms for the aquarium hobby due to its hardiness and suitability for community tanks. ''X. hellerii'' is a common laboratory animal and has been used as a
model organism A model organism (often shortened to model) is a non-human species that is extensively studied to understand particular biological phenomena, with the expectation that discoveries made in the model organism will provide insight into the workin ...
in studies involving female mate preference and male conflict. It is often designated ''X. helleri'' (with one 'i'), but authorities consider this an orthographic error and the spelling with two 'i's to be the valid specific epithet. It is named after Karl Bartholomaeus Heller, who collected the
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
. Due to interbreeding with the southern platyfish or 'platy', most swordtails in aquariums are hybrids to some degree. The males' elongated caudal fins have been found to significantly affect their chances at mating. The presence of a well-endowed male spurs the maturity of females, while it inhibits the maturity of juvenile males in the vicinity of the well-endowed male.


Breeding

Swordtails are some of the easiest fish for amateur aquarist to breed, if the conditions of the tank are appropriate and it contains both males and females reproduction will occur without intervention. Often all female groups which have come from a mixed gender tank will be pregnant on arrival, creating a breeding population when the fry sexually mature at around three months of age. Later stages of pregnancy can be seen without much difficulty as a dark gravid spot will develop near the anal fin. Males can be distinguished from females by the presence of a Gonopodium, a modified anal fin used to impregnate the female during breeding. A ratio of one male to three or four females is recommended to ensure that individual females will not be overly harassed to breed. Plants and other structure in the aquarium will offer fry a place to hide after birth. This is important as newborn fry will be eaten by other aquarium inhabitants if they cannot find shelter.


Gallery

File:Swordtail cultivar.JPG, Cultivated form of ''X. hellerii'' File:Xiphophorus hellerii fry 01.jpg, Fry File:Xiphophorus hellerii red wagtail female 01.jpg File:Xiphophorus helleri Annanasschwerttaeger Maennchen.JPG


References


External links

* * * *
Xiphophorus maculatus (Spanish)Xiphophorus hellerii profile with photographs of various strains on Fishes N Inverts

Xiphophorus hellerii gallery with photographs of various strains on Fishes N Inverts
{{Taxonbar, from=Q272188 Xiphophorus Live-bearing fish Ovoviviparous fish Fish described in 1848 Fish of Mexico Taxa named by Johann Jakob Heckel