Sveltonectes Insolitus
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''Sveltonectes'' (meaning "agile swimmer" in Greek) is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
platypterygiine Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous (Kimmeridgian - Cenomanian) of Asia, Australia, Europe, North America and South America. Currentl ...
ophthalmosaurid Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous (Bajocian - Cenomanian) worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the ex ...
ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, altho ...
s known from Ul’yanovsk region, western
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
.


Etymology

''Sveltonectes'' was named by Valentin Fischer, Edwige Masure, Maxim S. Arkhangelsky, and
Pascal Godefroit Pascal Godefroit is a Belgian paleontologist. He discovered dinosaurs like ''Olorotitan'' in 2003. Godefroit is the director of earth and life sciences at the Royal Belgian Institute of Natural Sciences The Museum of Natural Sciences of Belgium ...
in
2011 File:2011 Events Collage.png, From top left, clockwise: a protester partaking in Occupy Wall Street heralds the beginning of the Occupy movement; protests against Libyan dictator Muammar Gaddafi, who was killed that October; a young man celebrate ...
and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Sveltonectes insolitus''. The generic name is derived from ''sveltos'',
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
for "agile", and ''nektes'', Greek for "swimmer", and refers to its small size, streamlined skull, and powerful girdle musculature. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
is derived from ''insolitus'',
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "unusual", in reference to the numerous unusual features of this ichthyosaur, as well as its unusual tooth morphology.


Description

''Sveltonectes'' is known from the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
IRSNB R269, an almost complete three-dimensionally preserved
skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...
including a partial
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
. It was collected in unnamed locality, in Ul’yanovsk, dating to the late
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
stage of the
early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
, about 126
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
. Another
thunnosaur Thunnosauria (Greek for "tuna lizard" – ''thunnos'' meaning "tuna" and ''sauros'' meaning "lizard") is an extinct clade of parvipelvian ichthyosaurs from the Early Jurassic to the early Late Cretaceous (Hettangian–Cenomanian) of Asia, Austral ...
described from the same locality, by Bogolubow (1909), is ''"
Ichthyosaurus ''Ichthyosaurus'' (derived from Greek ' () meaning 'fish' and ' () meaning 'lizard') is a genus of ichthyosaurs from the Early Jurassic ( Hettangian - Pliensbachian), with possible Late Triassic record, from Europe (Belgium, England, Germany, S ...
" steleodon''. It is a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
'' that is twice the size and more robust than ''Sveltonectes''. Like some other ophthalmosaurids, ''Sveltonectes'' has a projection on the
nasal bone The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Eac ...
of the skull into the naris, a prefrontal forming part of the margin of the naris and a frontal forming part of the margin of the supratemporal fenestra. ''Sveltonectes'' is unique in that it has small, sharp teeth and numerous other peculiar features such as a very primitive prootic. The distinctive shape of these teeth suggest that it had a different feeding habit than other cretaceous ophthalmosaurids. Within the Ophthalmosauridae, ''Sveltonectes'' is most closely related to ''
Aegirosaurus ''Aegirosaurus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from the late Jurassic and early Cretaceous of Europe. It was originally named as a species of ''Ichthyosaurus''. Discovery and species Originally describe ...
''.


Phylogeny

The following cladogram shows a possible phylogenetic position of ''Sveltonectes'' in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020).


See also

*
List of ichthyosaurs This list of ichthyosaurs is a comprehensive listing of all genera that have ever been included in the order Ichthyosauria or the parent clade Ichthyopterygia, excluding purely vernacular terms. The list includes all commonly accepted genera, but a ...
*
Timeline of ichthyosaur research This timeline of ichthyosaur research is a chronological listing of events in the history of paleontology focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, shar ...


References

Early Cretaceous ichthyosaurs Fossil taxa described in 2011 Extinct animals of Russia Ophthalmosauridae Ichthyosauromorph genera {{cretaceous-reptile-stub