Sugar Gum
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''Eucalyptus cladocalyx'', commonly known as sugar gum, is a species of eucalypt tree found in the
Australian state The states and territories are federated administrative divisions in Australia, ruled by regional governments that constitute the second level of governance between the federal government and local governments. States are self-governing ...
of
South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
. It is found naturally in three distinct populations - in the Flinders Ranges,
Eyre Peninsula The Eyre Peninsula is a triangular peninsula in South Australia. It is bounded by the Spencer Gulf on the east, the Great Australian Bight on the west, and the Gawler Ranges to the north. Originally called Eyre’s Peninsula, it was named aft ...
and on Kangaroo Island.


Description

The tree notable for its mottled colourful yellow to orange bark, strongly discolourous leaves and
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s grouped on leafless branchlets inside the
tree crown The crown of a plant refers to the total of an individual plant's aboveground parts, including stems, leaves, and reproductive structures. A plant community canopy consists of one or more plant crowns growing in a given area. The crown of a wo ...
. The old
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...
is smooth and grey, shedding in irregular patches to expose the fresh yellowy-brown bark. Flowers are creamy-white in summer. The capsules are barrel to urn shaped. Sugar gums from the Flinders Ranges reach up to in height and have the classic "gum" habit - with a straight trunk having a diameter at breast height (dbh) of and steep branches occurring about halfway up. Each main branch ends with its own little canopy. They are commonly cultivated as farm
windbreak A windbreak (shelterbelt) is a planting usually made up of one or more rows of trees or shrubs planted in such a manner as to provide shelter from the wind and to protect soil from erosion. They are commonly planted in hedgerows around the edges ...
s and for timber. However, Eyre Peninsula and Kangaroo Island trees are much shorter, typically between in height, and often have crooked trunks and a dbh of . The crown has an open spreading habit with a typical spread of . The strongly discolorous, glossy adult leaves are arranged alternately supported on a petiole that is in length. The leaf blade is darker green on upper side and paler below with slightly falcate to
lanceolate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
shape, a length of and a width of , with a base usually tapering to the petiole. The side-veins in the leaf are at an acute or wider angle and densely reticulate. The intramarginal vein is parallel to but removed from margin, with small and obscure oil glands. It flowers in summer, producing white-cream-yellow flowers. The axillary unbranched
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
occur in groups of buds 7, 9 or 11 buds per umbel. The oblong pale green, yellow to creamy mature buds have a length of and a width of . The buds are often longitudinally striated and scarred, with a rounded operculum, inflexed stamens and cuboid to oblong anthers. The urceolate or barrel-shaped, longitudinally ribbed fruits that form after flowering are in length and wide, with a descending disc and three or four enclosed valves. The light grey to brown seeds within the fruit have a flattened-ovoid shape that can be pointed at one end and are long.


Taxonomy

The species was first formally described by the
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
Ferdinand von Mueller Baron Sir Ferdinand Jacob Heinrich von Mueller, (german: Müller; 30 June 1825 – 10 October 1896) was a German-Australian physician, geographer, and most notably, a botanist. He was appointed government botanist for the then colony of Vict ...
in 1853 in the journal ''Linnaea: Ein Journal für die Botanik in ihrem ganzen Umfange, oder Beiträge zur Pflanzenkunde''. In 1860, von Mueller referred to ''Eucalyptus corynocalyx'' in '' Fragmenta phytographiae Australiae'', citing several earlier publications, including the ''Linnaea'' journal article, but all only have a description of ''E. cladocalyx''. ''Eucalyptus corynocalyx'' is therefore a nomen illegitimum and a
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
of ''E. cladocalyx''. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
is taken from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
words ''klados'', meaning ''branch'', "twig" or "stem" and ''kalyx'', meaning "cup", "cover" or "outer envelope of a flower", in reference to the leafless branchlets that bear the flowers. In 2013,
Dean Nicolle 270px, Dean Nicolle and '' Eucalyptus deanei'' Dean Nicolle (born 1974), is an Australian botanist, arborist and ecologist. He is widely recognised as the leading authority on the genus ''Eucalyptus''. Nicolle was born in Adelaide, South Austr ...
describe three subspecies of ''E. cladocalyx'' and the names have been accepted by the
Australian Plant Census The Australian Plant Census (APC) provides an online interface to currently accepted, published, scientific names of the vascular flora of Australia, as one of the output interfaces of the national government Integrated Biodiversity Information Syst ...
: * ''Eucalyptus cladocalyx'' F.Muell. subsp. ''cladocalyx'' is a lower-growing, more spreading tree than the other subspecies and has short, broad leaves and larger fruit; * ''Eucalyptus cladocalyx'' subsp. ''crassa'' D.Nicolle is the tallest-growing subspecies and has long, narrow leaves and large fruit; * ''Eucalyptus cladocalyx'' subsp. ''petila'' D.Nicolle is a tall subspecies with erect branches, narrow adult leaves and relatively small fruit.


Distribution

''Eucalyptus cladocalyx'' is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to southern parts of South Australia. It is found in three distinct populations: in the southern and central-eastern parts of the Eyre Peninsula, through much of the Flinders Ranges, and on Kangaroo Island. It is most likely part of relic forests of wetter climates from the past. ''E. cladocalx'' has now become naturalised in the
South West The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...
region of Western Australia, in southern Victoria and, beyond its native range, in some parts of south-eastern South Australia. It is also naturalised overseas in northern and southern Africa, in
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,
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,
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and
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. Subspecies ''cladocalyx'' is restricted to the southern and eastern Eyre Peninsula, subspecies ''crassa'' to Kangaroo Island and subspecies ''petila'' to the southern Flinders Ranges.


Uses

The tree has been widely planted across southern Australia, often as a windbreak or shelterbelt, but also for timber and firewood production. The wood is
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
resistant, with moderate strength and durability, and can be used for furniture, flooring, posts, construction timber and railway sleepers. It is a fast-growing tree but is best planted in open sun in clay, loam or sandy soils. It is an efficient user of water and
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
and
frost Frost is a thin layer of ice on a solid surface, which forms from water vapor in an above-freezing atmosphere coming in contact with a solid surface whose temperature is below freezing, and resulting in a phase change from water vapor (a gas) ...
tolerant, with flowers that attract bees. It is also known to be a suitable breeding habitat for the
Yellow-tailed Black-Cockatoo The yellow-tailed black cockatoo (''Zanda funerea'') is a large cockatoo native to the south-east of Australia measuring in length. It has a short crest on the top of its head. Its plumage is mostly brownish black and it has prominent yellow ...
. ''E. cladocalyx'' is well adapted to regular bushfires and can resprout
epicormic An epicormic shoot is a shoot growing from an epicormic bud, which lies underneath the bark of a trunk, stem, or branch of a plant. Epicormic buds lie dormant beneath the bark, their growth suppressed by hormones from active shoots higher up ...
ally. It also produces a large number of seedlings through wind dispersal of seeds. The hard and heavy heartwood is a pale yellow-brown colour and has fine uniform texture with an interlocked grain. The density of air-dried wood is around and is moderately durable. The dwarf subspecies ''cladocalyx'' is sold in the nursery trade as ''E. cladocalyx'' 'Nana'.


Weed potential

''E. cladocalyx'' has invaded bushland in Western Australian, where it has been introduced. It has the capacity to spread up to 70 metres away from locations where it has been planted, and seems to survive bush fires more effectively than several local eucalypt species. It has become an invasive species in South Africa, where it is now registered as a category 2 invader plant.


References

* Holliday, I. ''A field guide to Australian trees (3rd edition)'', Reed New Holland, 2002 * Cronin, L. ''Key Guide to Australian Trees'', Envirobook, 2000
Rawlings, M. ''Regional allozyme divergence in Sugar Gum, Eucalyptus cladocalyx'', 2005, Centre for Plant Biodiversity Research
{{DEFAULTSORT:Eucalyptus cladocalyx Trees of Australia Trees of Mediterranean climate cladocalyx Flora of South Australia Myrtales of Australia Plants described in 1853 Taxa named by Ferdinand von Mueller