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''Suaeda nigra'', often still known by the former name ''Suaeda moquinii'', is a species of flowering plant in the amaranth family, known by the
vernacular name A vernacular or vernacular language is in contrast with a "standard language". It refers to the language or dialect that is spoken by people that are inhabiting a particular country or region. The vernacular is typically the native language, n ...
s bush seepweed or Mojave sea-blite.


Taxonomy

''Suaeda nigra'' was first formally described as a new species by
Constantine Samuel Rafinesque Constantine Samuel Rafinesque-Schmaltz (; October 22, 1783September 18, 1840) was a French 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ultimat ...
in 1832. Its
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
was collected by Edwin James along the
Canadian River The Canadian River is the longest tributary of the Arkansas River in the United States. It is about long, starting in Colorado and traveling through New Mexico, the Texas Panhandle, and Oklahoma. The drainage area is about .Texas panhandle The Texas Panhandle is a region of the U.S. state of Texas consisting of the northernmost 26 counties in the state. The panhandle is a square-shaped area bordered by New Mexico to the west and Oklahoma to the north and east. It is adjacent to ...
in 1820. In 1827
John Torrey John Torrey (August 15, 1796 – March 10, 1873) was an American botany, botanist, chemist, and physician. Throughout much of his career, he was a teacher of chemistry, often at multiple universities, while he also pursued botanical work, focus ...
tentatively misidentified this specimen as "''Chenopodium maritimum'' L. ?", but only in 1856 did Torrey finally describe the
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
as ''Chenopodina moquini''. In 1889
Edward Lee Greene Edward Lee Greene (August 20, 1843–November 10, 1915) was an American botanist known for his numerous publications including the two-part ''Landmarks of Botanical History'' and the describing of over 4,400 species of plants in the American W ...
moved it to the genus ''
Suaeda __NOTOC__ ''Suaeda'' is a genus of plants also known as seepweeds and sea-blites. Most species are confined to saline or alkaline soil habitats, such as coastal salt-flats and tidal wetlands. Many species have thick, succulent leaves, a character ...
'' (he continued to misspell it as ''moquini'').
Aven Nelson Aven Nelson (March 24, 1859 – March 31, 1952) was an American botanist who specialized in plants of the Rocky Mountains. He was one of the founding professors of the University of Wyoming, where he taught for 55 years as professor and served as ...
corrected the name to ''moquinii'' in 1909, and the species was often known under the name ''Suaeda moquinii'' until the 21st century. Rafinesque had named the Texas specimen ''Chenopodium nigrum'' on the basis of Torrey's 1827 summary description of the specimen. In 1918
James Francis Macbride James Francis Macbride (19 May 1892 16 June 1976) was an American botanist who devoted most of his professional life to the study of the flora of Peru. Early life and education Born on 19 May 1892 in Rock Valley, Iowa, Macbride graduated from th ...
moved it to the genus ''
Suaeda __NOTOC__ ''Suaeda'' is a genus of plants also known as seepweeds and sea-blites. Most species are confined to saline or alkaline soil habitats, such as coastal salt-flats and tidal wetlands. Many species have thick, succulent leaves, a character ...
'', but in 1977 C. O. Hopkins and W. H. Blackwell argued that this name was both a ''
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
'' and superfluous (not based on a real holotype), only for H. J. Schenk and W. R. Ferren Jr. to argue in 2001 that Rafinesque had clearly referenced Torrey's description of the specimen, and that it was thus provided with both a formal taxonomic description as well as a type.


Description

''Suaeda nigra'' is genetically diverse and quite variable in appearance. The species is a
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
or subshrub growing from a woody base with many spreading branches, reaching up to in height. The plants may be facultative
annual Annual may refer to: * Annual publication, periodical publications appearing regularly once per year **Yearbook ** Literary annual * Annual plant * Annual report * Annual giving * Annual, Morocco, a settlement in northeastern Morocco * Annuals (b ...
s when they find themselves growing in seasonally flooded wetlands. It may be hairy to hairless but is usually waxy. It may be green to red to dark purple in color, sometimes almost black. The
succulent In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meani ...
leaves are linear to lance-shaped, 1 to 3 centimetres long, and flat or cylindrical. Flowers occur in clusters along the upper stems, each cluster containing 1 to 12 flowers. Leaf-like
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s accompany the clusters. The flower has no petals and is composed of a calyx of fleshy, rounded
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s. The fruit is an utricle that grows within the calyx.


Distribution

''Suaeda nigra'' is native to much of western North America, from central Canada through the
Western United States The Western United States (also called the American West, the Far West, and the West) is the region comprising the westernmost states of the United States. As American settlement in the U.S. expanded westward, the meaning of the term ''the Wes ...
,
Great Basin The Great Basin is the largest area of contiguous endorheic basin, endorheic watersheds, those with no outlets, in North America. It spans nearly all of Nevada, much of Utah, and portions of California, Idaho, Oregon, Wyoming, and Baja California ...
, the
Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...
in
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
, and into northern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
.


Ecology

It grows in many types of
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
with saline and
alkaline In chemistry, an alkali (; from ar, القلوي, al-qaly, lit=ashes of the saltwort) is a base (chemistry), basic, ionic compound, ionic salt (chemistry), salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as ...
substrates, such as desert flats,
dry lake A dry lake bed, also known as a playa, is a basin or depression that formerly contained a standing surface water body, which disappears when evaporation processes exceeds recharge. If the floor of a dry lake is covered by deposits of alkaline c ...
s (locally called 'playas') and seeps. It is mostly an inland species but is occasionally seen in coastal areas, such as
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...
.Jepson
/ref>


References


External links


Jepson Manual Treatment of ''Suaeda nigra''''Suaeda nigra'' — U.C. Photo gallery
{{Taxonbar, from=Q3733922 nigra Halophytes Flora of the Western United States Flora of Western Canada Flora of Arizona Flora of California Flora of Colorado Flora of Idaho Flora of Nevada Flora of New Mexico Flora of Texas Flora of Utah Flora of Wyoming Flora of the California desert regions Flora of the Great Basin Flora of the Sierra Nevada (United States) Natural history of the California chaparral and woodlands Natural history of the Mojave Desert Barilla plants Flora without expected TNC conservation status