Sthenurinae (from ''Sthenurus'',

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

for 'strong-tailed') is a subfamily within the marsupial family

Macropodidae Macropodidae is a family of marsupials that includes kangaroos, wallabies, tree-kangaroos, wallaroos, pademelons, quokkas, and several other groups. These genera are allied to the suborder Macropodiformes, containing other macropods, and ar ...

, known as 'short faced kangaroos'. No members of this subfamily are extant today, with all becoming extinct by the late

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

. ''

Procoptodon goliah ''Procoptodon'' is an extinct genus of giant short-faced ( sthenurine) kangaroos that lived in Australia during the Pleistocene Epoch. ''P. goliah'', the largest known kangaroo species that ever existed, stood at about . They weighed about . Othe ...

'', the largest macropodid known to have existed, was a sthenurine kangaroo, but sthenurines occurred in a range of sizes, with '' Procoptodon gilli'' being the smallest at the size of a small wallaby. The short, robust skull of sthenurines is considered to be indicative that they were browsers that fed on leaves. Some species may have been able to reach above their heads and grasp branches with their semiopposable paws to assist in procuring leaves from trees. A single

hoof The hoof (plural: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits, yet the rumin ...

ed digit is present on the feet of sthenurines.

Taxonomy

The subfamilial arrangement Sthenurinae was circumscribed by

Ludwig Glauert Ludwig Glauert MBE (5 May 1879 – 1 February 1963) was a British-born Australian paleontologist, herpetologist and museum curator. He is known for work on Pleistocene mammal fossils, and as a museum curator who played an important role in na ...

in 1926.

Locomotion

Unlike modern macropodids, which hop (either bipedally or quadrupedally), sthenurines seem to have abandoned saltation as a means of locomotion. Their comparatively inflexible spines, robust hindlimb and pelvic elements, and the lack of capacity for rapid hopping suggest that these animals walked bipedally, somewhat like

hominids The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the east ...

, even converging with those primates in details of their pelvic anatomy. Furthermore, their hooved single digits and metatarsal anatomy suggest that unlike their

plantigrade 151px, Portion of a human skeleton, showing plantigrade habit In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. T ...

relatives, sthenurines were

digitigrade In terrestrial vertebrates, digitigrade () locomotion is walking or running on the toes (from the Latin ''digitus'', 'finger', and ''gradior'', 'walk'). A digitigrade animal is one that stands or walks with its toes (metatarsals) touching the groun ...

, walking on the tips of their "toes".

Taxonomy

Locomotion

References

Further reading