Staphylininae are a

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

rove beetle The rove beetles are a family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With roughly 63,000 species in thousands of genera, the ...

s (

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Staphylinidae). They contain the typical rove beetles with their long but fairly robust blunt-headed and -tipped bodies and short

elytra An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alterna ...

, as well as some more unusually-shaped lineages.

Systematics

As it seems, the Staphylininae are part of a large

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

together with the subfamilies Euaesthetinae,

Leptotyphlinae The Leptotyphlinae are a subfamily of the Staphylinidae, rove beetles.Newton, A. F., Jr., M. K. Thayer, J. S. Ashe, and D. S. Chandler. 2001. 22. Staphylinidae Latreille, 1802. p. 272–418. In: R. H. Arnett, Jr., and M. C. Thomas (eds.). Americ ...

, Megalopsidiinae, Oxyporinae,

Paederinae The Paederinae are a subfamily of the Staphylinidae, rove beetles. The Paederinae include two tribes, Paederini and Pinophilini. This insect is commonly known as Tomcat. Three of the genera of a subtribe of the Paederini are associated with ...

Pseudopsinae Pseudopsinae is a beetle subfamily of Staphylinidae The rove beetles are a family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. ...

, Scydmaeninae, Solieriinae,

Steninae Steninae is a subfamily of Staphylinidae.Newton, A. F., Jr., M. K. Thayer, J. S. Ashe, and D. S. Chandler. 2001. 22. Staphylinidae Latreille, 1802. p. 272–418. In: R. H. Arnett, Jr., and M. C. Thomas (eds.). American beetles, Volume 1. CRC Pres ...

, and the

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

Protactinae which are only known from

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s. The numerous Staphylininae

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...

are divided into six

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...

s, listed below along with some notable genera. However, a scientific study in 2020 proposed moving all tribes except Staphylinini to other subfamilies and raising several of Staphylinini's subtribes to tribal level. This would result in a new total of twelve tribes: Acylophorini, Afroquediini, Amblyopinini, Antimerini, †Baltognathini, Cyrtoquediini, Erichsoniini, Hyptiomini, Indoquediini, Quediini, Staphylinini and Tanygnathinini. * Diochini Casey, 1906 ** '' Antarctothius'' Coiffait & Saiz ** '' Coomania'' Cameron ** '' Diochus'' Erichson, 1839 *** '' Diochus electrus'' Chatzimanolis & Engel, 2011 (

Lutetian The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Midd ...

Baltic amber The Baltic region is home to the largest known deposit of amber, called Baltic amber or succinite. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that these forests created more than 1 ...

) * Maorothiini Diochus electrus ZooKeys-138-065-g001a

* Othiini C.G.Thomson, 1859 * Platyprosopini Lynch, 1884 * Staphylinini Latreille, 1802 (including Philonthini, Tanygnathinini) ** '' Cafius'' ** '' Creophilus'' *** '' Creophilus maxillosus'' – hairy rove beetle ** '' Eucibdelus'' ** '' Gabrius'' ** '' Jurecekia'' ** '' Ocypus'' *** ''

Ocypus olens The devil's coach-horse beetle (''Ocypus olens'') is a species of beetle belonging to the large family of the rove beetles (Staphylinidae). It was originally included in the genus '' Staphylinus'' in 1764, and some authors and biologists still us ...

'' – devil's coach-horse beetle *** '' Ocypus ophthalmicus'' ** '' Ontholestes'' ** '' Paederomimus'' ** '' Philonthus'' *** '' Philonthus cognatus'' ** '' Quedius'' *** '' Quedius curtipennis'' *** '' Quedius fuliginosus'' ** '' Rhynchocheilus'' ** '' Staphylinus'' ** '' Stevensia'' ** '' Thinopinus'' –

pictured rove beetle The pictured rove beetle (''Thinopinus pictus'') is a wingless rove beetle which lives on the sandy beaches of the West Coast of the United States from southern Alaska to Baja California. It is nocturnal, emerging at night from temporary sand bu ...

** ''

Velleius Marcus Velleius Paterculus (; c. 19 BC – c. AD 31) was a Roman historian, soldier and senator. His Roman history, written in a highly rhetorical style, covered the period from the end of the Trojan War to AD 30, but is most useful for the per ...

'' *** '' Velleius dilatatus'' * Xantholinini Erichson, 1839 ** '' Leptacinus'' *** '' Leptacinus intermedius'' ** '' Xantholinus''

References

External links

* * {{Staphylinidae-stub