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The Sri Lankan elephant (''Elephas maximus maximus'') is native to Sri Lanka and one of three recognised subspecies of the
Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living species of the genus ''Elephas'' and is distributed throughout the Indian subcontinent and Southeast Asia, from India in the west, Nepal in the no ...
. It is the type subspecies of the Asian elephant and was first described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...
under the
binomial Binomial may refer to: In mathematics *Binomial (polynomial), a polynomial with two terms * Binomial coefficient, numbers appearing in the expansions of powers of binomials *Binomial QMF, a perfect-reconstruction orthogonal wavelet decomposition ...
''Elephas maximus'' in 1758. The Sri Lankan elephant population is now largely restricted to the dry zone in the north, east and southeast of Sri Lanka. Elephants are present in
Udawalawe National Park Udawalawe National Park is a national park on the boundary of Sabaragamuwa and Uva Provinces in Sri Lanka. The park was created to provide a sanctuary for wild animals displaced by the construction of the Udawalawe Reservoir on the Walawe Rive ...
,
Yala National Park Yala (යාල) National Park is the most visited and second largest national park in Sri Lanka, bordering the Indian Ocean. The park consists of five blocks, two of which are now open to the public, and also adjoining parks. The blocks have ind ...
,
Lunugamvehera National Park Lunugamvehera National Park ( si, ලුණුගම්වෙහෙර ජාතික වනෝද්‍යානය) in Sri Lanka was declared in 1995, with the intention of protecting the catchment area of the Lunugamvehera reservoir and wildl ...
,
Wilpattu National Park Wilpattu National Park (''Willu-pattu'', "Land of Lakes") is a national park in Sri Lanka. The unique feature of this park is the existence of "Willus" (natural lakes) – natural, sand-rimmed water basins or depressions that fill with rainwater. ...
and
Minneriya National Park Minneriya National Park ( si, මින්නේරිය ජාතික වනෝද්‍යානය, translit=Minnēriya Jātika Vanōdyānaya; ta, மின்னேரியா தேசிய வனம், translit=Miṉṉēriyā Tēci ...
but also live outside protected areas. It is estimated that Sri Lanka has the highest density of elephants in Asia. Human-elephant conflict is increasing due to conversion of elephant habitat to settlements and permanent cultivation.


Characteristics

In general, Asian elephants are smaller than African elephants and have the highest body point on the head. The tip of their trunk has one finger-like process. Their back is convex or level. Females are usually smaller than males. Some males have
tusks Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share ...
. Sri Lankan elephants are the largest subspecies reaching a shoulder height of between , weigh between , and have 19 pairs of ribs. Their skin colour is darker than of '' indicus'' and of '' sumatranus'' with larger and more distinct patches of
depigmentation Depigmentation is the lightening of the skin or loss of pigment. Depigmentation of the skin can be caused by a number of local and systemic conditions. The pigment loss can be partial (injury to the skin) or complete (caused by vitiligo). It can be ...
on ears, face, trunk and belly. Only 7% of males bear tusks. Average adult elephant tusks grow up to about 6 feet. It can weight up to . Longest tusks of 7 feet 6 inches long was found in
Millangoda Raja Millangoda Raja (1938-30 July 2011: si, මිල්ලන්ගොඩ රාජා), also known as Millangoda tusker, was a Sri Lankan elephant. Over 9 feet tall and with 7 and a half foot long tusks, he was (wrongly) considered the longest tu ...
(1938-30 July 2011). The Sri Lankan subspecies designation is weakly supported by analysis of allozyme loci, but not by analysis of mitochondrial DNA (mtDNA) sequences. In July 2013, a
dwarf Dwarf or dwarves may refer to: Common uses *Dwarf (folklore), a being from Germanic mythology and folklore * Dwarf, a person or animal with dwarfism Arts, entertainment, and media Fictional entities * Dwarf (''Dungeons & Dragons''), a humanoid ...
Sri Lankan elephant was sighted in
Udawalawe National Park Udawalawe National Park is a national park on the boundary of Sabaragamuwa and Uva Provinces in Sri Lanka. The park was created to provide a sanctuary for wild animals displaced by the construction of the Udawalawe Reservoir on the Walawe Rive ...
. It was over tall but had shorter legs than usual and was the main aggressor in an encounter with a younger bull.


Distribution and habitat

Sri Lankan elephants are restricted mostly to the lowlands in the dry zone where they are still fairly widespread in north, south, east, north-western, north-central and south-eastern Sri Lanka. A small remnant population exists in the
Peak Wilderness Sanctuary Peak Wilderness sanctuary is a natural reserve in Sri Lanka. It is the third largest (by area) of the 50 sanctuaries in the country. "Sri Pada" Peak Wilderness sanctuary is a tropical rain forest that spreads over a land of 224 square kilometers ...
. They are absent from the wet zone of the country. Apart from Wilpattu and
Ruhuna National Park Yala (යාල) National Park is the most visited and second largest national park in Sri Lanka, bordering the Indian Ocean. The park consists of five blocks, two of which are now open to the public, and also adjoining parks. The blocks have i ...
s, all other
protected areas Protected areas or conservation areas are locations which receive protection because of their recognized natural, ecological or cultural values. There are several kinds of protected areas, which vary by level of protection depending on the ena ...
are less than in extent. Many areas are less than , and hence not large enough to encompass the entire home ranges of elephants that use them. In the Mahaweli Development Area, protected areas such as
Wasgomuwa National Park Wasgamuwa National Park is a natural park in Sri Lanka situated in the Matale and Polonnaruwa Districts. It was declared to protect and to make a refuge for the displaced wild animals during the Mahaweli Development Project in 1984 and is one of ...
,
Flood Plains National Park Flood Plains National Park ( si, ජලගැලුම් නිම්න ජාතික වනෝද්‍යානය, translit=Jalagælum Nimna Jātika Vanōdyānaya; ta, வெள்ளச் சமவெளிகள் தேசிய வ ...
,
Somawathiya National Park Somawathiya National Park is one of the four national parks designated under the Mahaweli River development project. Senarathna 2004: p. 166 Somawathiya Chaitya, a stupa said to be containing a relic of the tooth of the Buddha, is situated wit ...
and Trikonamadu Nature Reserve have been linked resulting in an overall area of of contiguous habitat for elephants. Nevertheless, about 65% of the elephant's range extends outside protected areas.


Former range

In the historical past, elephants were widely distributed from sea level to the highest mountain ranges. They occurred in the dry zone, in the lowland wet zone as well as in the cold damp montane forests. During the colonial period from 1505 to 1948, the wet zone was converted to commercially used fields and became heavily settled. Until 1830, elephants were so plentiful that their destruction was encouraged by the government, and rewards were paid for any that was killed. In the first half of the 19th century, forests in the montane zone were cleared large-scale for the planting of
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. Seeds of ...
, and afterward
tea Tea is an aromatic beverage prepared by pouring hot or boiling water over cured or fresh leaves of ''Camellia sinensis'', an evergreen shrub native to East Asia which probably originated in the borderlands of southwestern China and north ...
. The elephant population in the mountains was extirpated. During the
British rule The British Raj (; from Hindi ''rāj'': kingdom, realm, state, or empire) was the rule of the British Crown on the Indian subcontinent; * * it is also called Crown rule in India, * * * * or Direct rule in India, * Quote: "Mill, who was hims ...
, many bull elephants were killed by trophy hunters. One of the British army majors is credited with having shot over 1,500 elephants, and two others are reputed to have shot half that number each. Many other sportsmen have shot about 250–300 animals during this time. Between 1829 and 1855 alone, more than 6,000 elephants were captured and shot under order of colonial British Empire. By the turn of the 20th century, elephants were still distributed over much of the island. The area currently known as
Ruhuna National Park Yala (යාල) National Park is the most visited and second largest national park in Sri Lanka, bordering the Indian Ocean. The park consists of five blocks, two of which are now open to the public, and also adjoining parks. The blocks have i ...
was the ''Resident Sportsmen's Shooting Reserve'', an area reserved for the sporting pleasure of British residents in the country. In the early 20th century, mega reservoirs were constructed in the dry zone for irrigated agriculture. Ancient irrigation systems were rehabilitated and people resettled. This development gathered momentum after the independence in 1948. As a result, elephant habitat in the dry zone was severely fragmented.


Population trend

The size of wild elephant populations in Sri Lanka was estimated at * 19,500 in the early 19th century; * 10,000 in the early 20th century; * 7,000 to 8,000 in around 1920; * between 1,745 and 2,455 individuals in 1969; * between 2,500 and 3,435 in 1987; * 1,967 in June 1993 that were fragmented in five regions; * between 3,150 and 4,400 in 2000; * 3,150 in 2006; * 2,900–3,000 in 2007; * 5,879 in 2011, on the basis of counting elephants at water holes in the dry season. * 7,500 in 2019;


Ecology and behaviour

Elephants are classified as
megaherbivore Megaherbivores (Greek μέγας megas "large" and Latin ''herbivora'' "herbivore") are large terrestrial herbivores that can exceed in weight. This polyphyletic group of megafauna includes elephants, rhinos, hippos, and giraffes. The largest ...
s and consume up to of plant matter per day. As generalists, they feed on a wide variety of food plants. In Sri Lanka's northwestern region, feeding behaviour of elephants was observed during the period of January 1998 to December 1999. The elephants fed on a total of 116 plant species belonging to 35 families including 27 species of cultivated plants. More than half of the plants were non-tree species, i.e. shrub, herb, grass, or climbers. More than 25% of the plant species belonged to the family ''Leguminosae'', and 19% of the plant species belonged to the family of
true grasses Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
. The presence of cultivated plants in dung does not result solely due to raiding of crops as it was observed that elephants feed on leftover crop plants in fallow chenas. Juvenile elephants tend to feed predominantly on grass species. Food resources are abundant in regenerating forests, but at low density in mature forests. Traditional
slash-and-burn agriculture Slash-and-burn agriculture is a farming method that involves the cutting and burning of plants in a forest or woodland to create a field called a swidden. The method begins by cutting down the trees and woody plants in an area. The downed veget ...
creates optimum habitat for elephants through promoting successional vegetation. Females and calves generally form small, loosely associated social groups without the hierarchical tier structure exhibited by African savannah elephants. However, at some locations such as
Minneriya National Park Minneriya National Park ( si, මින්නේරිය ජාතික වනෝද්‍යානය, translit=Minnēriya Jātika Vanōdyānaya; ta, மின்னேரியா தேசிய வனம், translit=Miṉṉēriyā Tēci ...
, hundreds of individuals aggregate during the dry season, suggesting that grouping behaviour is flexible and depends on season and place. Like all
Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living species of the genus ''Elephas'' and is distributed throughout the Indian subcontinent and Southeast Asia, from India in the west, Nepal in the no ...
s, the Sri Lankan subspecies communicates using visual, acoustic and chemical signals. At least fourteen different vocal and acoustic signals have been described, which include some low-frequency calls that contain infrasonic frequencies.


Threats

During the armed conflict in Sri Lanka, elephants were maimed or killed by
land mine A land mine is an explosive device concealed under or on the ground and designed to destroy or disable enemy targets, ranging from combatants to vehicles and tanks, as they pass over or near it. Such a device is typically detonated automati ...
s. Between 1990 and 1994, a total of 261 wild elephants died either as a result of gunshot injuries, or were killed by poachers and land mines. Several elephants stepped on land mines and were crippled. Today, given the rarity of tuskers in Sri Lanka,
poaching Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...
for ivory is not a major threat. Nevertheless, some trade in ivory still goes on.
Kandy Kandy ( si, මහනුවර ''Mahanuwara'', ; ta, கண்டி Kandy, ) is a major city in Sri Lanka located in the Central Province. It was the last capital of the ancient kings' era of Sri Lanka. The city lies in the midst of hills ...
has been identified as the centre for such illegal trade. The greatest threat to elephants comes from an expanding human population and its demand for land. Loss of significant extents of elephant range to development continues currently, with a number of irrigation and development projects leading to the conversion of more elephant ranges to irrigated agriculture and settlements. Between 1999 and the end of 2006 every year nearly 100 wild elephants were killed. Elephants are killed to protect crops and houses. Other threats are poaching, deforestation, drought and starvation. During drought seasons many elephants damage agricultural land for food. Nearly 80 elephants were killed in north western Sri Lanka, 50 in south and east and another 30 in other parts of the country, totalling 160 elephant deaths in 2006 alone. Sri Lanka has thus become the country with the largest number of elephant fatality per year. DWC official recode showed that more than 361–405 elephants were killed in 2019.


Conservation

''Elephas maximus'' is listed on
CITES Appendix I CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of intern ...
. The elephant conservation strategy of the Department of Wildlife Conservation aims at conserving as many viable populations as possible in as wide a range of suitable habitats as is feasible. This means protecting elephants both within the system of protected areas and as many animals outside these areas that the land can support and landholders will accept, and not restricting elephants to the protected area network alone. * In the
Pinnawala Elephant Orphanage Pinnawala Elephant Orphanage ( si, පින්නවල අලි අනාථාගාරය), is an orphanage, nursery and captive breeding ground for wild Asian elephants located at Pinnawala village, northeast of Kegalle town in Sabara ...
in Kegalle injured elephants are treated, and orphaned baby elephants cared for. Nearly 70 elephants live here. Captive breeding is also going on. * The
Udawalawe Elephant Transit Centre The Udawalawe Elephant Transfer Home is a facility within Udawalawe National Park in Sri Lanka that was established in 1995 by the Sri Lanka Department of Wildlife Conservation. Its primary objective is to rehabilitate orphaned elephant calves ...
in
Udawalawe National Park Udawalawe National Park is a national park on the boundary of Sabaragamuwa and Uva Provinces in Sri Lanka. The park was created to provide a sanctuary for wild animals displaced by the construction of the Udawalawe Reservoir on the Walawe Rive ...
is a rehabilitation centre, where orphaned elephant calves are being kept until they can be released into the wild.


Cultural and symbolism

Elephants were a common element in
Sinhalese Sinhala may refer to: * Something of or related to the Sinhalese people of Sri Lanka * Sinhalese people * Sinhala language, one of the three official languages used in Sri Lanka * Sinhala script, a writing system for the Sinhala language ** Sinha ...
and Sri Lankan Tamils heraldry for over two thousand years and remained so through British colonial rule. The
coat of arms A coat of arms is a heraldic visual design on an escutcheon (i.e., shield), surcoat, or tabard (the latter two being outer garments). The coat of arms on an escutcheon forms the central element of the full heraldic achievement, which in its ...
and the flag of Ceylon Government from 1875 to 1948 included an elephant and even today many institutions use the Sri Lankan elephant in their coat of arms and insignia. An important cultural symbiosis has continued to exist between the elephant and humans for over two thousand years – no religious procession was complete without its retinue of elephants, and many large Buddhist temples and
Hindu Temple A Hindu temple, or ''mandir'' or ''koil'' in Indian languages, is a house, seat and body of divinity for Hindus. It is a structure designed to bring human beings and gods together through worship, sacrifice, and devotion.; Quote: "The Hin ...
s in Sri Lanka had their own elephants.


Captive elephants

Since time immemorial, elephants have been domesticated for uses as work elephants and
war elephant A war elephant was an elephant that was trained and guided by humans for combat. The war elephant's main use was to charge the enemy, break their ranks and instill terror and fear. Elephantry is a term for specific military units using elepha ...
s in Sri Lanka by the ancient kings. Elephants were exported from the island for hundreds of years and into the Portuguese and Dutch colonial era. The British did not export elephants, instead took to
hunting Hunting is the human practice of seeking, pursuing, capturing, or killing wildlife or feral animals. The most common reasons for humans to hunt are to harvest food (i.e. meat) and useful animal products ( fur/ hide, bone/tusks, horn/antler, ...
wild elephants and capture of wild for domestication as work elephants continued. Elephant Kraals were organised to capture large herds of elephants in the late nineteenth and early twentieth century. The capture of wild elephants were regulated under the Fauna and Flora Protection Ordinance in 1937, with the issuance of permits to capture of wild elephants. This practice stopped following the last Elephant Kraal in 1950 by Sir Francis Molamure. A census of the domesticated elephant population in 1970 indicated 532 elephants among 378 owners, while this number had dropped 344 in 1982. These domesticated elephants were used mainly as work elephants and for cultural pageants, the chief of which is the annual
Kandy Esala Perahera The Kandy Esala Perahera (the Sri Dalada Perahara procession of Kandy) also known as The Festival of the Tooth is a festival held in July and August in Kandy, Sri Lanka. This historical procession is held annually to pay homage to the Sacred T ...
. In recent years, the domesticated elephant population has dropped further with the need for their labour dropping widespread use of tractors. However, they remain in use in terrain inaccessible by vehicles for logging and used for tourism. Ownership of elephants are highly prestigious among Singhalese as a status symbol and calls have been made for permission to capture wild elephants or release of orphaned wild elephants in government care to Temples to take part in pageants. Captive breeding in private ownership does not take place due to the long period of unemployability associated with it. Legal reforms pertaining to the captive elephant population was introduced in 2021, just as a landmark case into dozens of calves being stolen from their herds in a ten-year period collapsed with the Attorney General's Department dropping charges and releasing the elephants to their former owners.


See also

* Sri Lankan ivories * List of mammals of Sri Lanka


References


External links

{{Taxonbar, from=Q583372 Endemic fauna of Sri Lanka Mammals of Sri Lanka Elephants Articles containing video clips