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''Vaccinium membranaceum'' is a species within the group of Vaccinium commonly referred to as
huckleberry Huckleberry is a name used in North America for several plants in the family Ericaceae, in two closely related genera: ''Vaccinium'' and ''Gaylussacia''. The huckleberry is the state fruit of Idaho. Nomenclature The name 'huckleberry' is a Nort ...
. This particular species is known by the common names thinleaf huckleberry, tall huckleberry, big huckleberry, mountain huckleberry, square-twig blueberry, and (ambiguously) as "
black huckleberry Black huckleberry is a common name for several plants and may refer to: *''Gaylussacia baccata'', native to eastern North America *''Vaccinium membranaceum ''Vaccinium membranaceum'' is a species within the group of Vaccinium commonly referred t ...
".


Distribution and habitat

''Vaccinium membranaceum'' is native to western North America, with a range extending in the northern from southern
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
,
Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
, and
Northwest Territories The Northwest Territories (abbreviated ''NT'' or ''NWT''; french: Territoires du Nord-Ouest, formerly ''North-Western Territory'' and ''North-West Territories'' and namely shortened as ''Northwest Territory'') is a federal territory of Canada. ...
south as far as
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
and the northern mountains of
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
. It can be found from the mountains next to the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
in the west, to the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
and
Black Hills The Black Hills ( lkt, Ȟe Sápa; chy, Moʼȯhta-voʼhonáaeva; hid, awaxaawi shiibisha) is an isolated mountain range rising from the Great Plains of North America in western South Dakota and extending into Wyoming, United States. Black Elk P ...
in the east. Isolated populations of this species have been found in
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
,
North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the Native Americans in the United States, indigenous Dakota people, Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north a ...
,
Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over to ...
, the
Upper Peninsula of Michigan The Upper Peninsula of Michigan – also known as Upper Michigan or colloquially the U.P. – is the northern and more elevated of the two major landmasses that make up the U.S. state of Michigan; it is separated from the Lower Peninsula by t ...
, and
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
. ''Vaccinium membranaceum'' grows at higher elevations in
subalpine Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
and
alpine Alpine may refer to any mountainous region. It may also refer to: Places Europe * Alps, a European mountain range ** Alpine states, which overlap with the European range Australia * Alpine, New South Wales, a Northern Village * Alpine National Pa ...
environments. It occurs in both
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
and
spruce A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
dominated forests and in open meadow ecosystems. In forests ''V. membranaceum'' often dominates the forest
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abov ...
during early to mid stages of
succession Succession is the act or process of following in order or sequence. Governance and politics *Order of succession, in politics, the ascension to power by one ruler, official, or monarch after the death, resignation, or removal from office of ...
.US Forest Service Fire Ecology
/ref> ''Vaccinium membranceum'' is fire adapted. The leaves and stems of the huckleberry are resistant to low-intensity fires, and if burned away they will resprout vigorously from rhizomes buried under the soil.


Description

''Vaccinium membranaceum'' is an erect
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
growing up to in maximum height. The new twigs are yellow-green and somewhat angled. The
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
leaves are alternately arranged. The very thin to membranous, oval leaf blades are up to 5 centimeters (2 inches) long. The edges are serrated, with each tiny tooth tipped with a glandular hair. Solitary flowers occur in the leaf axils. Each is around 6 millimeters (1/4 inch) long,
urn An urn is a vase, often with a cover, with a typically narrowed neck above a rounded body and a footed pedestal. Describing a vessel as an "urn", as opposed to a vase or other terms, generally reflects its use rather than any particular shape or ...
-shaped to cylindrical, and pale pink to waxy bronze in color. They are
pollinated Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, ...
by
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s. The mature fruit ranges from red through bluish-purple to a dark, almost black
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
about a centimeter wide. Each fruit contains an average of 47 tiny seeds. ;Reproduction The plant rarely reproduces via seed, rather, it usually spreads by
cloning Cloning is the process of producing individual organisms with identical or virtually identical DNA, either by natural or artificial means. In nature, some organisms produce clones through asexual reproduction. In the field of biotechnology, cl ...
itself from its
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
or shoots. The seeds do
germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
if dispersed by animals, however, as evidenced by populations of the plant growing on the recovering section of
Mount St. Helens Mount St. Helens (known as Lawetlat'la to the indigenous Cowlitz people, and Loowit or Louwala-Clough to the Klickitat) is an active stratovolcano located in Skamania County, Washington, in the Pacific Northwest region of the United St ...
. Other than the study by Yang et al. (2008) reports of ''V. membranaceum'' sprouting from seed are quite rare with other scientists who have studied this species reporting only 6 seedlings observed during 18 years in the field.


Uses

''Vaccinium membranaceum'' is the species that is the most commonly collected of all of the wild western huckleberries, and it has great commercial importance.Flora of North America, ''Vaccinium membranaceum'' Douglas ex Torrey, 1874. Thinleaf huckleberry
/ref> In a good year ''Vaccinium membranaceum'' shrubs produce a lot of fruit. The amount of fruit produced by these shrubs is legendary, with stories being told of mountain sides turned purple by all of the fruit, or shrubs being weighed to the ground by large, and abundant berries


Native Americans

Both humans and wildlife enjoy feasting on this fruit in the late summer and early fall. People have been eating the fruit of this species for thousands of years. It was and continues to be widely used for food by Native Americans. The
Kutenai The Kutenai ( ), also known as the Ktunaxa ( ; ), Ksanka ( ), Kootenay (in Canada) and Kootenai (in the United States), are an indigenous people of Canada and the United States. Kutenai bands live in southeastern British Columbia, northern ...
called the black huckleberry shawíash (
Ktunaxa The Kutenai ( ), also known as the Ktunaxa ( ; ), Ksanka ( ), Kootenay (in Canada) and Kootenai (in the United States), are an indigenous people of Canada and the United States. Kutenai bands live in southeastern British Columbia, northern ...
: ǂawiyaǂ).
Alaska Natives Alaska Natives (also known as Alaskan Natives, Native Alaskans, Indigenous Alaskans, Aboriginal Alaskans or First Alaskans) are the indigenous peoples of Alaska and include Iñupiat, Yupik, Aleut, Eyak, Tlingit, Haida, Tsimshian, and a numbe ...
consumed it in bread and pies as a source of
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) an ...
, the
Coeur d'Alene people The Coeur d'Alene (also ''Skitswish''; natively ''Schi̲tsu'umsh'') are a Native American nation and one of five federally recognized tribes in the state of Idaho. The Coeur d'Alene have sovereign control of their Coeur d'Alene Reservation, w ...
ate the fruit fresh, dried, mashed, cooked, and added it to soup or froze it for later use, and many other groups relished it and stored it frozen, dried, pressed into cakes, or canned for winter use.


Wildlife

The plant also provides a key food source for
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white have o ...
and
grizzly bear The grizzly bear (''Ursus arctos horribilis''), also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America. In addition to the mainland grizzly (''Ursus arctos horri ...
s, which eat the leaves, stems, roots, and fruit.
Elk The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The common ...
,
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
, and
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
also browse the plant. The thickets provide cover for many species of small animals.


Symbol

The huckleberry is the official state fruit of
Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...
, with this particular species assumed to be the huckleberry in question.


Management

Some Native American groups lit carefully planned
controlled burn A controlled or prescribed burn, also known as hazard reduction burning, backfire, swailing, or a burn-off, is a fire set intentionally for purposes of forest management, farming, prairie restoration or greenhouse gas abatement. A control ...
s in wild huckleberry patches to promote fruit production by eliminating competing plants and by stimulating the huckleberry to sprout and spread. Native American groups throughout the
Pacific Northwest The Pacific Northwest (sometimes Cascadia, or simply abbreviated as PNW) is a geographic region in western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though ...
still utilize this plant as an important cultural food and are active in its management.Martin, L. P., et al. (2008)
Management and monitoring plan for the enhancement of big huckleberry in Government Meadows, Mt Baker-Snoqualmie National Forest.
USDA Forest Service.


See also

*''
Gaylussacia baccata ''Gaylussacia baccata'', the black huckleberry, is a common huckleberry found throughout a wide area of eastern North America. Distribution The plant is native to Eastern Canada and the Great Lakes region, the Midwestern and Northeastern Unit ...
'' — which shares the common name "black huckleberry"


References


External links


Jepson Manual Treatment of ''Vaccinium membranaceum''United States Department of Agriculture Plants Profile for ''Vaccinium membranaceum''Washington Burke Museum, University of Washington''Vaccinium membranaceum'' — Calphotos Photo gallery, University of Californiaphoto of herbarium specimen at Missouri Botanical Garden, collected in Oregon in 2006
{{Taxonbar, from=Q292442 membranaceum Berries Flora of Western Canada Plants described in 1874 Plants used in Native American cuisine Flora of the Northwestern United States Flora of the Southwestern United States Flora of South Dakota Flora of Michigan Flora of Ontario Flora without expected TNC conservation status