''Squalius'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...

found in

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

Hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...

ization is not rare in the Cyprinidae, including this genus. '' S. alburnoides'' is known to be of ancient hybrid origin, with the

paternal A father is the male parent of a child. Besides the paternal bonds of a father to his children, the father may have a parental, legal, and social relationship with the child that carries with it certain rights and obligations. An adoptive fathe ...

lineage deriving from a

prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...

species related to ''

Anaecypris ''Anaecypris hispanica'', the Spanish minnowcarp, is a small species of ray-finned fish species in the family Cyprinidae. It is the only living member of the genus ''Anaecypris''. It is endemic to the Iberian Peninsula and is found in the basin o ...

''; the latter mated with ancestral '' S. pyrenaicus''. Present-day ''S. alburnoides'' mates with

sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species sh ...

congeners of other

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

.Collares-Pereira, M.J. & Coelho, M.M. (2010)
Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex ''Squalius alburnoides''.
''Journal of Fish Biology, 76 (3): 707–715.''

Species

There are currently 52 recognized species in this genus: * ''

Squalius adanaensis The Adana chub (''Squalius adanaensis'') is a cyprinid fish endemic to the lower Seyhan basin in Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country lo ...

Turan Turan ( ae, Tūiriiānəm, pal, Tūrān; fa, توران, Turân, , "The Land of Tur") is a historical region in Central Asia. The term is of Iranian origin and may refer to a particular prehistoric human settlement, a historic geographical re ...

Kottelat Maurice Kottelat (born 16 July 1957 in Delémont, SwitzerlandC ...

Doğan Doğan is both a masculine Turkish given name and a Turkish surname meaning ''Falcon''. Notable people with the name include: Given name * Doğan Abukay, Turkish experimental physicist and academic * Doğan Akhanlı (1957–2021) Turkish-German w ...

, 2013 (Adana chub) Turan, D., Kottelat, M. & Doğan, E. (2013)
Two new species of ''Squalius'', ''S. adanaensis'' and ''S. seyhanensis'' (Teleostei: Cyprinidae), from the Seyhan River in Turkey.
''Zootaxa, 3637 (3): 308–324.'' * ''

Squalius agdamicus ''Squalius agdamicus'' is a cyprinid fish endemic to the lower Kura basin in Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transconti ...

'' S. N. Kamensky, 1901 * ''

Squalius alburnoides ''Squalius alburnoides'' is a species of fish in the family Cyprinidae. It is found in Portugal and Spain. Its natural habitats are rivers and intermittent rivers. It may be threatened by habitat loss. This species is a highly peculiar fish in r ...

'' ( Steindachner, 1866) * ''

Squalius albus The Trasimeno chub (''Squalius albus'') is a cyprinid fish endemic to Lake Trasimeno in Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle o ...

'' ( Bonaparte, 1838) * ''

Squalius anatolicus ''Squalius anatolicus'' is a species of ray-finned fish in the family Cyprinidae. It is found only in Turkey. Its natural habitat is freshwater lake A lake is an area filled with water, localized in a basin, surrounded by land, and distin ...

'' ( Bogutskaya, 1997) (Beyşehir chub) * ''

Squalius aphipsi The Aphips chub (''Squalius aphipsi'') is a cyprinid fish endemic to the Kuban drainages in Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest c ...

'' (A. I. Aleksandrov, 1927) (Aphips chub) * ''

Squalius aradensis ''Squalius aradensis'' is a ray-finned fish species in the family Cyprinidae, growing to SL. It is endemic to Portugal where it is found in only three river basins: Arade The Arade () is a river located in the region of the Algarve, souther ...

'' ( Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) * ''

Squalius aristotelis The Tuzla chub (''Squalius aristotelis'') is a cyprinid fish endemic to the Lake Tuzla drainage in Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country ...

'' Özuluğ & Freyhof, 2011 (Tuzla chub) Özuluğ, M. & Freyhof, J. (2011)
Revision of the genus ''Squalius'' in Western and Central Anatolia, with description of four new species (Teleostei: Cyprinidae).
''Ichthyological Exploration of Freshwaters, 22 (2): 107-148.'' * '' Squalius berak'' Heckel, 1843 (Mesopotamian chub) Esmaeili, H.R., Khaefi, R. & Zamanian Nejad, R. (2016)
Historical review on the taxonomy of ''Squalius berak'' Heckel, 1843 (Teleostei: Cyprinidae).
''FishTaxa, 1 (3): 118-126.'' * '' Squalius cappadocicus'' Özuluğ & Freyhof, 2011 (Cappadocian chub) * '' Squalius carinus'' Özuluğ & Freyhof, 2011 (Chocolate chub) * ''

Squalius carolitertii ''Squalius carolitertii'' is a species of fresh-water fish in the family Cyprinidae. It is found in Portugal and Spain, and known there as the ''bordallo, escalo'' or ''gallego''. Its natural habitats are rivers and intermittent rivers. It is ...

'' ( Doadrio, 1988) * '' Squalius castellanus'' Doadrio,

Perea Perea or Peraea (Greek: Περαία, " the country beyond") was the portion of the kingdom of Herod the Great occupying the eastern side of the Jordan River valley, from about one third the way down the Jordan River segment connecting the Sea ...

& F. M. Alonso, 2007 * '' Squalius cephaloides'' ( Battalgil, 1942) (Thick-lipped chub) * ''

Squalius cephalus ''Squalius cephalus'' is a European species of freshwater fish in the carp family Cyprinidae. It frequents both slow and moderate rivers, as well as canals and still waters of various kinds. This species is referred to as the common chub, Europ ...

'' (

Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

, 1758) (Common chub) * '' Squalius cii'' ( J. Richardson, 1857) * '' Squalius fellowesii'' ( Günther, 1868) (Aegean chub) * '' Squalius ghigii'' ( Gianferrari, 1927) (Rhodes chub) * ''

Squalius illyricus ''Squalius illyricus'', the Illyrian chub, is a species of ray-finned fish in the family Cyprinidae. It inhabits karstic waters of Bosnia and Herzegovina, Croatia and Albania. Its natural habitats are rivers and water storage areas. It is threat ...

'' Heckel & Kner, 1857 * '' Squalius janae'' Bogutskaya & Zupančič, 2010 * '' Squalius keadicus'' ( Stephanidis, 1971) * '' Squalius kosswigi'' ( M. S. Karaman ( sr), 1972) * '' Squalius kottelati''

Yılmaz Yılmaz () is a Turkish word that translates to "unshirking", "unbeatable", or "brave", and is a very common surname and occasional male given name. Given name * Yekta Yılmaz Gül (born 1978), Turkish Greco-Roman wrestler * Yılmaz Arslan (born ...

Kaya Kaya may refer to: People *Kaya (given name) *Kaya (surname) Places *Kaya, Burkina Faso, a town in Burkina Faso, capital of the department *Kaya Airport, serving the town * Kaya Department, a department or commune of Sanmatenga Province in centr ...

, 2009 * '' Squalius laietanus'' Doadrio,

Kottelat Maurice Kottelat (born 16 July 1957 in Delémont, SwitzerlandC ...

& de Sostoa, 2007 * '' Squalius lepidus'' Heckel, 1843 * ''

Squalius lucumonis ''Squalius lucumonis'' is a species of freshwater fish in the family Cyprinidae which is found only in Italy. Its natural habitat is within rivers A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, ...

'' (

Bianco Bianco is a town and ''comune'' in the Province of Reggio Calabria, in southern Italy. It is a seaside town and a popular tourist resort. The main attractions are the remainings of an old abbey and the ruins of a Roman house. See also *Calabri ...

, 1983) * '' Squalius malacitanus'' Doadrio & Carmona, 2006 * '' Squalius microlepis'' Heckel, 1843 * '' Squalius moreoticus'' ( Stephanidis, 1971) (Stymphalia chub) * '' Squalius namak'' Khaefi, Esmaeili, Sayyadzadeh, Geiger & Freyhof, 2016 Khaefi, R., Esmaeili, H.R., Sayyadzadeh, G., Geiger, M.F. & Freyhof, J. (2016): ''Squalius namak'', a new chub from Lake Namak basin in Iran (Teleostei: Cyprinidae). ''Zootaxa, 4169 (1): 145–159.'' * '' Squalius orientalis'' ( Nordmann, 1840) * '' Squalius orpheus''

Kottelat Maurice Kottelat (born 16 July 1957 in Delémont, SwitzerlandC ...

& Economidis, 2006 (Orpheus chub) * '' Squalius pamvoticus'' ( Stephanidis, 1939) (Pamvotis chub) * '' Squalius peloponensis'' (

Valenciennes Valenciennes (, also , , ; nl, label=also Dutch, Valencijn; pcd, Valincyinnes or ; la, Valentianae) is a commune in the Nord department, Hauts-de-France, France. It lies on the Scheldt () river. Although the city and region experienced a s ...

, 1844) (Peloponnese chub) * '' Squalius platyceps'' Zupančič, Marić, Naseka & Bogutskaya, 2010 * ''

Squalius prespensis ''Squalius prespensis'' is a species of freshwater fish in the family Cyprinidae, growing to SL. It was originally described as a subspecies of European chub. ''Squalius prespensis'' is endemic to the Lake Prespa basin, shared between Albani ...

'' ( Fowler, 1977) * '' Squalius pursakensis'' ( Hankó ( hu), 1925) (Sakarya chub) * ''

Squalius pyrenaicus ''Squalius pyrenaicus'' is a species of freshwater fish in the family Cyprinidae. It is found in Portugal and Spain. Its natural habitats are rivers, intermittent rivers, and water storage areas. It is becoming rare due to habitat loss Ha ...

'' ( Günther, 1868) * '' Squalius recurvirostris'' Özuluğ & Freyhof, 2011 (Akşehir chub) * '' Squalius seyhanensis''

Kottelat Maurice Kottelat (born 16 July 1957 in Delémont, SwitzerlandC ...

, 2013 (Seyhan chub) * '' Squalius spurius'' Heckel, 1843 (Orontes chub) * '' Squalius squaliusculus''

Kessler Kessler or Keßler (in German) may refer to: * Kessler (automobile), an American automobile made 1921–1922 * Kessler (name), people named Kessler * ''Kessler'' (TV series), a British television series from 1981 * Kessler, Ohio, an unincorporat ...

, 1872 (Syr-Darya dace) * '' Squalius squalus'' ( Bonaparte, 1837) * '' Squalius svallize'' Heckel & Kner, 1857 * '' Squalius tenellus'' Heckel, 1843 * '' Squalius torgalensis'' ( Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998) * '' Squalius turcicus'' De Filippi, 1865 (Transcaucasian chub) * '' Squalius ulanus'' Günther, 1899 * '' Squalius valentinus'' Doadrio & Carmona, 2006 * '' Squalius vardarensis'' S. L. Karaman, 1928 * '' Squalius zrmanjae'' S. L. Karaman, 1928

References

{{Taxonbar, from=Q2725287 Cyprinidae genera Taxa named by Charles Lucien Bonaparte Taxonomy articles created by Polbot