The spruce grouse (''Canachites canadensis''), also known as Canada grouse, spruce hen or fool hen,
is a medium-sized
grouse
Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondria ...
closely associated with the coniferous
boreal forest
Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...
s or
taiga
Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...
of North America. It is the only member of the
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Canachites''.
It is one of the most
arboreal
Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
grouse species, fairly well adapted to perching and moving about in trees. When approached by a predator, it relies on camouflage and immobility to an amazing degree; for example, letting people approach within a few feet before finally taking flight.
Taxonomy
In 1747 the English naturalist
George Edwards included an illustration and a description of a female spruce grouse in the second volume of his ''A Natural History of Uncommon Birds''. He used the English name "The Brown and Spotted Heathcock". Edwards based his hand-coloured etching on a preserved specimen that had been sent to him in London from the
Hudson Bay
Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
in Canada by a Mr Light. Edwards was later sent what he assumed was a specimen of the male bird by
James Isham James Isham (1716–1761) was chief factor (master) at both York Factory and Fort Prince of Wales in Canada during the mid-1700s. He kept detailed journals that described life in the region, including flora and fauna that were unknown to people in ...
. In 1750 he included the male bird in the third volume of his book under the English name "The Black and Spotted Heathcock". When in 1758 the Swedish naturalist
Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
updated his ''
Systema Naturae
' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' for the
tenth edition, he placed the spruce grouse with other grouse in the
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Tetrao
''Tetrao'' is a genus of birds in the grouse subfamily known as capercaillies. They are some of the largest living grouse.
Taxonomy
The genus ''Tetrao'' was introduced in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his ...
''. Linnaeus included a brief description, coined the
binomial name
In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Tetrao canadensis'' and cited Edwards' work. The spruce grouse is now the only species placed in the genus ''Canachites'' that was introduced in 1885 by the zoologist
Leonhard Stejneger
Leonhard Hess Stejneger (30 October 1851 – 28 February 1943) was a Norwegian-born American ornithologist, herpetologist and zoologist. Stejneger specialized in vertebrate natural history studies. He gained his greatest reputation with reptiles ...
.
The genus name ''Canachites'' combines the name ''
Canace
In Greek mythology, Canace (; Ancient Greek: Κανάκη means "‘barking") was a Thessalian princess as daughter of King Aeolus of Aeolia and Enarete, daughter of Deimachus. She was sometimes referrred as Aeolis.
Family
Canace was the sister ...
'' from
Greek mythology
A major branch of classical mythology, Greek mythology is the body of myths originally told by the Ancient Greece, ancient Greeks, and a genre of Ancient Greek folklore. These stories concern the Cosmogony, origin and Cosmology#Metaphysical co ...
with the
Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''-itēs'' meaning "resembling".
In the first half of the 20th century, spruce grouse were classified as two separate species in the genus ''Canachites'': spruce grouse (''C. canadensis'') and
Franklin's grouse (''C. franklinii''), a position reinstated by Birdlife International in 2014. However, as of early 2021 the
International Ornithological Congress
International is an adjective (also used as a noun) meaning "between nations".
International may also refer to:
Music Albums
* ''International'' (Kevin Michael album), 2011
* ''International'' (New Order album), 2002
* ''International'' (The T ...
(IOC)
, the
American Ornithological Society
The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
, and the
Clements taxonomy
''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world.
The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...
retain Franklin's grouse as a subspecies of spruce grouse.
The species was later moved to the genus ''Dendragapus'', congeneric with the
blue grouse
The genus ''Dendragapus'' contains two closely related species of grouse that have often been treated as a single variable taxon (blue grouse). The two species are the dusky grouse (''Dendragapus obscurus'') and the sooty grouse (''Dendragapus f ...
with which spruce grouse often share coniferous habitats. However, spruce grouse do not have inflatable cervical sacs as in blue grouse, and the natal plumage of the two species is different. Based on its stronger resemblance to the
Siberian grouse
The Siberian grouse (''Falcipennis falcipennis''), also known as Siberian spruce grouse, Amur grouse, or Asian spruce grouse, is a short, rotund forest-dwelling grouse. A sedentary, non-migratory bird, it is the only member of the genus ''Falcipe ...
(''Falcipennis falcipennis''), the spruce grouse was later reclassified into the genus ''Falcipennis''. However, taxonomic studies found this classification to be
paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
, with the Siberian grouse being
basal to a clade containing the spruce grouse, ''
Tetrao
''Tetrao'' is a genus of birds in the grouse subfamily known as capercaillies. They are some of the largest living grouse.
Taxonomy
The genus ''Tetrao'' was introduced in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his ...
'', and ''
Lyrurus
''Lyrurus'' is a genus of birds in the grouse subfamily. They are known as black grouse because the male's plumage of both species is colored black as its base colour.
Taxonomy
The genus ''Lyrurus'' was introduced in 1832 by the English natural ...
''. Due to this, the spruce grouse was again reclassified in ''Canachites'', bringing its taxonomy full-circle. Six subspecies are recognized by the IOC in these ranges:
[
* ''C. c. osgoodi'' Bishop, 1900 – north Alaska
* ''C. c. atratus'' Grinnell, 1910 – south Alaska
* ''C. c. canadensis'' (Linnaeus, 1758) – central Alberta to Labrador and Nova Scotia. Introduced in ]Newfoundland
Newfoundland and Labrador (; french: Terre-Neuve-et-Labrador; frequently abbreviated as NL) is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region ...
in 1964 and Anticosti Island
; moe, Notiskuan; mic, Natigostec
, sobriquet =
, image_name = RiviereHuileAnticosti.jpg
, image_caption = Salmon fisherman on Rivière à l'Huile
, image_map ...
in 1985–86.
* ''C. c. canace'' (Linnaeus, 1766) – southeast Canada and northeast United States
* ''C. c. isleibi'' (Dickerman & Gustafson, 1996) – southeast Alaska
* ''C. c. franklinii'' (Douglas
Douglas may refer to:
People
* Douglas (given name)
* Douglas (surname)
Animals
*Douglas (parrot), macaw that starred as the parrot ''Rosalinda'' in Pippi Longstocking
*Douglas the camel, a camel in the Confederate Army in the American Civil W ...
, 1829) – southeast Alaska to northwest Wyoming and Idaho
Description
Spruce grouse are long; males weigh and females . The spruce grouse has a wingspan range of 21.5-22.6 in (54.5-57.5 cm). Races vary slightly in plumage, especially in the tail pattern and in the extent of white on the underparts, but in general adult males are mainly grey above and black below, with white spots along the side, and a red patch of bare skin over the eye called the "eyebrow comb". This red eyebrow comb, should not be confused by a similar yellow marking found on other species of grouse including, but not limited to, sharp-tailed and dusky grouse
The dusky grouse (''Dendragapus obscurus'') is a species of forest-dwelling grouse native to the Rocky Mountains in North America.del Hoyo, J., Elliott, A., & Sargatal, J., eds. (1994). ''Handbook of the Birds of the World'' 2: 401-402. Lynx Edi ...
. Adult females are mottled brown (red morph) or mottled grey (grey morph) with dark and white bars on the underparts. Juveniles resemble females. Females may be confused with ruffed grouse
The ruffed grouse (''Bonasa umbellus'') is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska. It is the most widely distributed game bird in North America. It is non-migratory. It is the only specie ...
but they have a dark tail with a pale band at the end (while the reverse is true in ruffed grouse) and they do not erect their crown feathers when alarmed the way ruffed grouse do.
Spruce grouse are among the most silent of all grouse, but they nevertheless have a number of calls used to warn of predators, to repel territorial intruders, to maintain brood cohesion, or to elicit brooding. In the subspecies ''franklinii'', territorial males are notable for their wing-clap display. At the end of a short flight through the trees, the wings are brought together over the back so as to produce two sharp claps, about 0.5 s apart, loud enough to be heard by the human ear 150 m away. These territorial displays can sometimes be elicited by similar hand clapping, and can be used to detect male territories and estimate their density. Other non-vocal sounds include soft drumming by territorial males (a sound produced by the beating wings) and a tail-swish and a whoosh produced by the tail feathers in flight.
Distribution and habitat
As a specialist of the taiga
Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...
, the spruce grouse is found throughout Canada. In the United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
, it is present in Alaska, northern New England
New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
, the Adirondacks
The Adirondack Mountains (; a-də-RÄN-dak) form a massif in northeastern New York with boundaries that correspond roughly to those of Adirondack Park. They cover about 5,000 square miles (13,000 km2). The mountains form a roughly circular d ...
in New York
New York most commonly refers to:
* New York City, the most populous city in the United States, located in the state of New York
* New York (state), a state in the northeastern United States
New York may also refer to:
Film and television
* '' ...
, northern Michigan
Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and the ...
, northeastern Minnesota
Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over to ...
, northern Wisconsin
Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...
and the montane conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous forests of Montana
Montana () is a state in the Mountain West division of the Western United States. It is bordered by Idaho to the west, North Dakota and South Dakota to the east, Wyoming to the south, and the Canadian provinces of Alberta, British Columbi ...
, Idaho
Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...
, Maine
Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and north ...
, Oregon
Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
, and Washington
Washington commonly refers to:
* Washington (state), United States
* Washington, D.C., the capital of the United States
** A metonym for the federal government of the United States
** Washington metropolitan area, the metropolitan area centered o ...
. Spruce grouse are always associated with conifer-dominated forests, be they pine
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
, spruce
A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
, or fir. They seem to prefer young successional stands. In summer they can be found near rich understory of blueberries
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' within the genus ''Vaccinium''. ''Vaccinium'' also includes cranberries, bi ...
and other shrub, and in winter they prefer denser stands.
The spruce grouse is essentially a permanent resident. However, in southwestern Alberta up to 35% of females and 7% of males move several kilometers (up to 11) between summer breeding sites and wintering sites. Similar movements have also been documented in Alaska and New Brunswick (Canada).
Behaviour and ecology
This species prefers to walk on the ground or along tree limbs rather than fly. Like other grouse, in the fall they grow "snow shoes" (short lateral extensions, or pectinations) on their toes. This increases the surface area of the toes and helps support the bird on snow and probably to grip branches as well. The pectinations are shed in the spring.
Flights are usually over short distances, most commonly from the ground to a tree nearby, or vice versa. Flight can be rapid but no actual measures of velocity have been made.
Breeding
Males are promiscuous; they disperse and advertise a territory that is visited by females for mating. Females are solely responsible for the rest of the reproductive effort though males have been known to stay with young chicks and seem to aid in keeping the brood together. For a nest, they scratch a depression in the ground in a bush or under a low-lying coniferous branch or fallen tree, away from other females and from the males' territories. The nest is lined with grasses, leaves, and a few feathers. Nesting season is from early May to early July. Up to 10 eggs may be laid, the usual number being 4–7. Laying rate is 1 egg every 1.4 days. Eggs are about and are tawny olive or buff, marked with blotches of brown. Incubation begins with the last egg laid and lasts about 24 days. Young are about 15 g at hatching (0.5 ounces) and they are precocial
In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
; they walk out of the nest as soon as they are dry (about 8 h after hatching). They are capable of fluttering up from the ground at 1 week of age. The brood stays together and is accompanied by the hen, who broods them all night and frequently during the day until the young are 3–5 weeks old. Brooding behaviour of the female seems to be initiated by specific calls from the chicks when they are cold. At 70–100 days of age, chicks tend to leave the group and become independent.
Females breed only once a year. Most females first breed at 1 year of age, but about half the males delay establishing a territory until 2 years. The species' lifespan appears to be about 5–6 years in the wild, though one study in Southwestern Alberta has found two males and one female that lived to be at least 13 years old.[Boag, D.A. and M.A. Schroeder (1992). Spruce Grouse. In ''The Birds of North America, No.5'' (A, Poole, P. Stettenheim, and F. Gill, eds.), Philadelphia: The Academy of Natural Sciences; Washington, DC: The American Ornithologists' Union.]
Food and feeding
The staple winter food is conifer needles, clipped directly from the tree, preferably the midcrown of pines though other conifers such as spruce are exploited as well. Spruce needles are high in calcium and their increase in use by females in Spring may be related to egg production In summer the birds can forage on the ground, eating berries
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
, green plants such as blueberry leaves, fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, and some insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s.[ In winter, when only needles are consumed, the caeca (dead-end extensions of the intestines) and ventriculus (gizzard) increase in size to support digestion. The ]crop
A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydroponic ...
is also well developed: up to 45 cc of needles (about 10% of body mass) can be stored in the crop at the end of the day, to be digested over the duration of the night fast. Like other birds, spruce grouse consume clay, grit or small stones to help their gizzard
The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including archosaurs (pterosaurs, crocodiles, alligators, dinosaurs, birds), earthworms, some gastropods, so ...
break down food. Chicks under 1 week old feed on insects and other arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s, then switch to berries and fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
until the fall, when they start feeding on needles. Birds captured while on the winter diet have been maintained several months without loss of weight, on only pine needles and grit and clay taken from where they were seen consuming it.
Predators
Spruce grouse eggs are taken by American red squirrel
The American red squirrel (''Tamiasciurus hudsonicus'') is one of three species of tree squirrels currently classified in the genus ''Tamiasciurus'', known as the pine squirrels (the others are the Douglas squirrel, ''T. douglasii'', and the sou ...
s, gray fox
The gray fox (''Urocyon cinereoargenteus''), or grey fox, is an omnivorous mammal of the family Canidae, widespread throughout North America and Central America. This species and its only congener, the diminutive island fox (''Urocyon littora ...
es, weasel
Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets and European mink. Members of this genus are small, active predators, with long and slender bo ...
s, and possibly corvid
Corvidae is a Cosmopolitan distribution, cosmopolitan family (biology), family of oscine passerine birds that contains the crows, ravens, rook (bird), rooks, jackdaws, jays, magpies, treepies, choughs, and nutcracker (bird), nutcrackers. In coll ...
s. Adults can fall prey to various hawk
Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica.
* The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
s and owl
Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s, red fox
The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
es, American
American(s) may refer to:
* American, something of, from, or related to the United States of America, commonly known as the "United States" or "America"
** Americans, citizens and nationals of the United States of America
** American ancestry, pe ...
and Pacific marten
The Pacific marten (''Martes caurina'') is a species of North American mammal, a member of the family Mustelidae. It is found throughout western North America.
Taxonomy
The species was formerly considered conspecific with the American marten ( ...
s, wolves
The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
, cougar
The cougar (''Puma concolor'') is a large Felidae, cat native to the Americas. Its Species distribution, range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mamm ...
s and coyote
The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
s.
Relationship with humans
This species is commonly hunted. Annual bag figures in the late 1970s were about 188,000 birds in the US and about 360,000 in Canada.[ Even though it has been deemed of Least Concern by IUCN, in the United States this bird is a protected species according to many northern states' hunting rules. The spruce grouse is available for hunting in Alaska from August 10 - March 31 and in some regions through May 15th according to the Alaska Department of Fish and Game.
Conversely, the spruce grouse is considered threatened in its southern range - being listed as ]Endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
in New York and Vermont
Vermont () is a state in the northeast New England region of the United States. Vermont is bordered by the states of Massachusetts to the south, New Hampshire to the east, and New York to the west, and the Canadian province of Quebec to ...
, and ''Critically Imperiled'' in Wisconsin according to NatureServe
NatureServe, Inc. is a non-profit organization based in Arlington County, Virginia, US, that provides proprietary wildlife conservation-related data, tools, and services to private and government clients, partner organizations, and the public. Nat ...
. In New York, the population is estimated at 175-315 individuals, with declines recorded since the 1970s. However, a management plan has been developed.
Gallery
File:Tetras-du-Canada.jpg, alt=Female spruce grouse, Grands-Jardins National Park, Female spruce grouse, Grands-Jardins National Park, Québec
File:Spruce Grouse (Falcipennis canadenis) RWD.jpg, Male, Acadia National Park, Maine
File:Sgrouse1.jpg, Male spruce grouse during mating season
File:Spruce Grouse Chick.jpg, A chick, Gros Morne National Park, Newfoundland
References
* del Hoyo, J., A. Elliott, and J. Sargatal (eds.) (1994). ''Handbook of the Birds of the World.'' Volume 2. Barcelona: Lynx Edicions.
* Ehrlich, P.R., D.S. Dobkin, and D. Wheye (1988). ''The Birder's Handbook: A Field Guide to the Natural History of North American Birds.'' New York: Simon and Schuster Inc.
* Terres, J.K. (1982). ''The Audubon Society Encyclopedia of North American Birds.'' New York: Alfred A. Knopf.
*Pendergast, B.A. and D.A. Boag (1971). "Maintenance and Breeding of spruce Grouse in Captivity" Journal of Wildlife Management 35(1):177-179.
*Pendergast, B.A. and D.A. Boag (1970). “Seasonal Changes in Diet of Spruce Grouse in Central Alberta” Journal of Wildlife Management 34(3):605-609.
*Pendergast, B.A. and D.A. Boag (1973). “Seasonal Changes in the Internal Anatomy of Spruce Grouse in Alberta” The Auk Vol. 90,(2): 307- 317.
*Pendergast, B.A. and D.A. Boag (1971). "Nutritional Aspects of the Diet of Spruce Grouse in Central Alberta" The Condor Vol. 73, (4): 437- 443.
External links
Spruce Grouse Factsheet
New York Department of Environmental Conservation
Spruce Grouse
Cornell Lab of Ornithology
{{Taxonbar, from=Q19056
spruce grouse
The spruce grouse (''Canachites canadensis''), also known as Canada grouse, spruce hen or fool hen, is a medium-sized grouse closely associated with the coniferous boreal forests or taiga of North America. It is the only member of the genus ''C ...
spruce grouse
The spruce grouse (''Canachites canadensis''), also known as Canada grouse, spruce hen or fool hen, is a medium-sized grouse closely associated with the coniferous boreal forests or taiga of North America. It is the only member of the genus ''C ...
Native birds of Alaska
Birds of Canada
spruce grouse
The spruce grouse (''Canachites canadensis''), also known as Canada grouse, spruce hen or fool hen, is a medium-sized grouse closely associated with the coniferous boreal forests or taiga of North America. It is the only member of the genus ''C ...
spruce grouse
The spruce grouse (''Canachites canadensis''), also known as Canada grouse, spruce hen or fool hen, is a medium-sized grouse closely associated with the coniferous boreal forests or taiga of North America. It is the only member of the genus ''C ...
Taxobox binomials not recognized by IUCN