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The thrush nightingale (''Luscinia luscinia''), also known as the sprosser, is a small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
that was formerly classed as a member of the thrush family Turdidae, but is now more generally considered to be an
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
, Muscicapidae. It, and similar small
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
an species, are often called chats. It is a migratory insectivorous species breeding in forests in Europe and the
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
and overwintering in Africa. The distribution is more northerly than the very closely related
common nightingale The common nightingale, rufous nightingale or simply nightingale (''Luscinia megarhynchos''), is a small passerine bird best known for its powerful and beautiful song. It was formerly classed as a member of the thrush family Turdidae, but is no ...
, ''Luscinia megarhynchos'', which it closely resembles in appearance. It nests near the ground in dense undergrowth. The thrush nightingale is similar in size to the
European robin The European robin (''Erithacus rubecula''), known simply as the robin or robin redbreast in Great Britain & Ireland, is a small insectivorous passerine bird that belongs to the chat subfamily of the Old World flycatcher family. About in len ...
. It is plain greyish-brown above and white and greyish-brown below. Its greyer tones, giving a cloudy appearance to the underside, and lack of the
common nightingale The common nightingale, rufous nightingale or simply nightingale (''Luscinia megarhynchos''), is a small passerine bird best known for its powerful and beautiful song. It was formerly classed as a member of the thrush family Turdidae, but is no ...
's obvious rufous tail side patches are the clearest plumage differences from that species. Sexes are similar. It has a similar but more powerful song than that of the nightingale.


Etymology

"Nightingale" is derived from "night", and the
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
''galan'', "to sing". The genus name ''Luscinia'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for the common nightingale.


Description

An adult thrush nightingale is about long with a wingspan of approximately . The head, nape and the whole of the upper parts of the thrush nightingale are dark brown with a slight olive tinge. The colour is much deeper than that of the nightingale and is not at all rufous. The upper tail-coverts are less olivaceous and the tail feathers are dark rufous-brown. The lores and ear-coverts are brownish-black and the chin and throat are pale buff or whitish, mottled with brown, and are paler in colour than the nightingale. The sides of the throat are spotted brown and the pale feathers of the breast have brown central bands giving the breast a mottled appearance. The under tail-coverts are buff, sometimes barred or marked with brown. The wing feathers and wing-coverts are dark brown and less rufous than the nightingale. The beak, legs and feet are brown and the irises are dark brown. The sexes are similar to each other in appearance and the juveniles are darker and more mottled. There is a single
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
in July and August at the end of the breeding season.


Voice

The male's is loud, with a range of whistles, trills and clicks and includes a flute-like "''pioo''" with a pure bell-like tone. It is sometimes interrupted by a rasping "''dserr''" sound and is rather solemn as compared to that of the nightingale. The song does not have that bird's loud whistling crescendo and is quite distinctive. It is also sometimes sung in the bird's winter quarters. The call-note "''whit''" resembles that of the nightingale but is higher pitched and more abrupt.


Distribution and habitat

The thrush nightingale is a migrant species. It breeds in eastern Europe and the western part of temperate Asia. The northern limit of its summer range extends to
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...
, southern
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of B ...
,
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...
and
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
, the
Baltic States The Baltic states, et, Balti riigid or the Baltic countries is a geopolitical term, which currently is used to group three countries: Estonia, Latvia, and Lithuania. All three countries are members of NATO, the European Union, the Eurozone, ...
, the
Republic of Karelia The Republic of Karelia (russian: Респу́блика Каре́лия, Respublika Kareliya; ; krl, Karjalan tašavalta; ; fi, Karjalan tasavalta; vep, Karjalan Tazovaldkund, Ludic: ''Kard’alan tazavald''), also known as just Karelia (rus ...
,
Kostroma Kostroma ( rus, Кострома́, p=kəstrɐˈma) is a historic city and the administrative center of Kostroma Oblast, Russia. A part of the Golden Ring of Russian cities, it is located at the confluence of the rivers Volga and Kostroma. Popu ...
,
Vologda Vologda ( rus, Вологда, p=ˈvoləɡdə) is a types of inhabited localities in Russia, city and the administrative center of Vologda Oblast, Russia, located on the river Vologda (river), Vologda within the watershed of the Northern Dvina. ...
,
Perm Perm or PERM may refer to: Places *Perm, Russia, a city in Russia ** Permsky District, the district **Perm Krai, a federal subject of Russia since 2005 **Perm Oblast, a former federal subject of Russia 1938–2005 **Perm Governorate, an administra ...
,
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
,
Turkestan Turkestan, also spelled Turkistan ( fa, ترکستان, Torkestân, lit=Land of the Turks), is a historical region in Central Asia corresponding to the regions of Transoxiana and Xinjiang. Overview Known as Turan to the Persians, western Turke ...
and Altai. The southern limit extends from
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
and the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
, through
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
,
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...
, southern
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
,
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
, the
Crimea Crimea, crh, Къырым, Qırım, grc, Κιμμερία / Ταυρική, translit=Kimmería / Taurikḗ ( ) is a peninsula in Ukraine, on the northern coast of the Black Sea, that has been occupied by Russia since 2014. It has a pop ...
and northern
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...
. It overwinters in Africa south of the
Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
. It is an occasional visitor to the British Isles. In its breeding range, the thrush nightingale is found in damp deciduous woodland typically with
alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...
and
birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...
. It favours thick undergrowth with
bramble A bramble is any rough, tangled, prickly shrub, usually in the genus ''Rubus'', which grows blackberries, raspberries, or dewberries. "Bramble" is also used to describe other prickly shrubs, such as roses (''Rosa'' species). The fruits inclu ...
s, dense shrubs and tangled vegetation in swampy places and near water. In its winter quarters it is found in dense patches of thorn bush, especially in valley bottoms near water courses, and sometimes in thick vegetation at the edge of woodland. The bird is a host of the
acanthocephala Acanthocephala (Greek , ', thorn + , ', head) is a phylum of parasitic worms known as acanthocephalans, thorny-headed worms, or spiny-headed worms, characterized by the presence of an eversible proboscis, armed with spines, which it uses to p ...
n intestinal parasite '' Apororhynchus silesiacus''.


Behaviour

The thrush nightingale feeds chiefly on the ground taking
earthworm An earthworm is a terrestrial invertebrate that belongs to the phylum Annelida. They exhibit a tube-within-a-tube body plan; they are externally segmented with corresponding internal segmentation; and they usually have setae on all segments. Th ...
s,
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s and the adults,
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e and
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
e of insects such as
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s, small
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s,
ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
s and
flies Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
. In the autumn, the berries of currants (''Ribes'' spp.) and elders (''Sambucus'' spp.) are also eaten. Before crossing the
Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
on its migration, thrush nightingales build up their fat reserves. It has been found experimentally that magnetic cues may stimulate the birds to do this. A simulation of the magnetic field found in northern Egypt encouraged birds preparing to migrate from Sweden to further build up their body fat. The thrush nightingale breeds in damp forests, nesting on the ground, often in the middle of a bed of
stinging nettle ''Urtica dioica'', often known as common nettle, burn nettle, stinging nettle (although not all plants of this species sting) or nettle leaf, or just a nettle or stinger, is a herbaceous perennial flowering plant in the family Urticaceae. Or ...
s (''Urtica dioica''). The nest rests on a platform of dead leaves and is composed of dead grass stalks, bents (''
Agrostis ''Agrostis'' (bent or bentgrass) is a large and very nearly cosmopolitan genus of plants in the grass family, found in nearly all the countries in the world. It has been bred as a GMO creeping bent grass. Species * '' Agrostis aequivalvi'' ...
'' spp.), sedges and stems, lined with finer material. It is built by the female which lays four or five (occasionally six) eggs. These are a milky-blue colour, usually plain but sometimes with a slight speckling of rusty-brown and measure an average of . The hen incubates the eggs which hatch in about thirteen days. The young are fed by both parents and
fledge Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
when about eleven days old, but are not fully independent for another twelve days or so.


Status

BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
estimates that there are between 11 and 20 million thrush nightingales in Europe and that, as Europe forms somewhere between 50% and 74% of the bird's global range, the total world population may be between 15 and 41 million individuals. In Europe, the population seems to be increasing slightly. The bird is considered to be of ''
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
'' by the International Union for Conservation of Nature
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
.


References


External links


Avibase
* Thrush nightingale
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q206130
thrush nightingale The thrush nightingale (''Luscinia luscinia''), also known as the sprosser, is a small passerine bird that was formerly classed as a member of the thrush family Turdidae, but is now more generally considered to be an Old World flycatcher, Muscic ...
Birds of Europe Birds of Russia Birds of Central Africa Birds of East Africa Birds of North Africa
thrush nightingale The thrush nightingale (''Luscinia luscinia''), also known as the sprosser, is a small passerine bird that was formerly classed as a member of the thrush family Turdidae, but is now more generally considered to be an Old World flycatcher, Muscic ...
thrush nightingale The thrush nightingale (''Luscinia luscinia''), also known as the sprosser, is a small passerine bird that was formerly classed as a member of the thrush family Turdidae, but is now more generally considered to be an Old World flycatcher, Muscic ...