Spotted Scat
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''Scatophagus argus'', the spotted scat, butterfish, mia mia, spotted butterfish or tiger scat, is a species of fish in the scat family Scatophagidae. It occurs in two basic color morphs which are called green scat and ruby or red scat. This fish is generally distributed around the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region, to Japan, New Guinea, and southeastern Australia. They live in coastal muddy areas, including estuaries, mangroves, harbours, and the lower courses of rivers. They are popular aquarium fish.


Taxonomy

''Scatophagus argus'' was first formally described in 1766 as ''Chaetodon argus'' by Carl Linnaeus with the
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
given as India. In 1831 Georges Cuvier described the genus '' Scatophagus'' and Linnaeus's ''C. argus'' was designated as its type species. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''argus'' refers to the mythical hundred-eyed guardian of Io,
Argus Argus is the Latinized form of the Ancient Greek word ''Argos''. It may refer to: Greek mythology * See Argus (Greek myth) for mythological characters named Argus **Argus (king of Argos), son of Zeus (or Phoroneus) and Niobe **Argus (son of Ar ...
, who following his death had his eyes became the feathers of a peacock, a reference to the brown to reddish-brown spots on the body of this fish.


Description

''Scatophagus argus'' has a body which is rectangular and strongly compressed with the head having a steep dorsal profile. It has a moderately large eye which has a diameter noticeably smaller than the length of the rounded, snout. They have a small, horizontal mouth which is not protractile. There are a number of rows of small bristle like teeth in the jaws. The dorsal fin has 10-11 spines and 16-18 soft rays, while the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
has 4 spines and 13-15 soft rays. Spines and rays of the dorsal fin are separated by a deep notch and the first spine in the dorsal fin lies flat. The rear margins of soft parts of the dorsal and anal fins is roughly vertical. The caudal fin is rounded in juveniles and truncate to weakly emarginate adults. Small ctenoid scales cover the body. The body is greenish-brown to silvery with many brown to red-brown spots. Juveniles are a greenish-brown with either a few large, dark, rounded blotches, or five or six dark, vertical bars. This species attains a maximum total length of .


Distribution and habitat

''Scatophagus argus'' has a wide Indo-Pacific range. It is found from the Persian Gulf, along the south Asian coast into the western Pacific. It occurs as far north as Japan south to New South Wales,
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, and Fiji. It has also been recorded from
French Polynesia )Territorial motto: ( en, "Great Tahiti of the Golden Haze") , anthem = , song_type = Regional anthem , song = " Ia Ora 'O Tahiti Nui" , image_map = French Polynesia on the globe (French Polynesia centered).svg , map_alt = Location of Frenc ...
. A small population was probably established in the Mediterranean Sea around Malta after a first report in 2007 (probably as a result of released aquarium fishes). This is a species of sheltered, shallow coastal waters such as estuaries, harbours, mangrove pools, and the lower parts of fresh water streams, particularly where there are high mineral concentrations. The very small juveniles float within the surface film of the water.


Biology

''Scatophagus argus'' is omnivorous and an indiscriminate eater. In 1992, biologists Barry and Fast reported adult scat from the Philippines were primarily herbivorous, while the juveniles preferred zooplankton. Although scat were named for their purported habit of feeding on offal, the name of the genus ''Scatophagus'' means "dung eater". it may be a misnomer as this behaviour has not been confirmed in diet studies. Since spotted scats can live in relatively enclosed waterbodies, as well as quite far upstream in freshwater rivers, they can adapt to varying salinities. As fry, they live in freshwater environments, but as they mature, they move to saltwater environments. They do not live in temperate waters, as they require at least a little warmth between This species forms schools. The females attain sexual maturity at around 7-9 months of age and when the weigh , whereas the males are sexually mature at a smaller size. In the Philippines, spawning is brought on by the monsoon rains that start in June and July and the increased rainfall brings cooler temperatures, increased river outflows and lower salinities. The eggs are about in diameter, they are transparent and have a spherical shape. The larvae take around 20 hours to hatch from fertilisation and on hatching are in length. The juveniles pass through a pelagic tholichthys larval stage, like butterflyfishes.


Utilisation

''Scatophagus argus'' is fished for and eaten by some people from its original environment, and can sting with small spikes in its anterior parts, inflicting a venom that causes great pain and dizziness. Treatment of the wound is often done by soaking the site of invenomation in hot water. It also appears in the
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
trade.


Parasites

''Scatophagus argus'' harbors parasites, like most fish. The acanthocephalan worm ''Pararhadinorhynchus magnus'' has been described from the intestine of this fish in waters off Vietnam.


References

{{DEFAULTSORT:Scatophagus Argus
argus Argus is the Latinized form of the Ancient Greek word ''Argos''. It may refer to: Greek mythology * See Argus (Greek myth) for mythological characters named Argus **Argus (king of Argos), son of Zeus (or Phoroneus) and Niobe **Argus (son of Ar ...
Fish described in 1766 Taxa named by Carl Linnaeus