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The spotted sandgrouse (''Pterocles senegallus'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of ground dwelling
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
in the family
Pteroclidae Sandgrouse is the common name for Pteroclidae , a family of sixteen species of bird, members of the order Pterocliformes . They are traditionally placed in two genera. The two central Asian species are classified as '' Syrrhaptes'' and the othe ...
. It is found in arid regions of northern and eastern Africa and across the Middle East and parts of Asia as far east as northwest India. It is a gregarious, diurnal bird and small flocks forage for seed and other vegetable matter on the ground, flying once a day to a waterhole for water. In the breeding season pairs nest apart from one another, the eggs being laid in a depression on the stony ground. The chicks leave the nest soon after hatching and eat dry seed, the water they need being provided by the male which saturates its belly feathers with water at the waterhole. The spotted sandgrouse is listed as being of "least concern" by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of nat ...
in its
Red List of Threatened Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
.


Description

The spotted sandgrouse reaches a length of about . The male has a small reddish-brown nape surrounded by a band of pale grey that extends to the bill and round the neck in a collar. The chin, neck and throat are orange and the breast grey. The upper parts are pinkish-grey with dark flight feathers and dark patches on the wings, tail and lower belly. The primaries are pale with dark trailing edges, a fact that distinguishes this species from the
crowned sandgrouse The crowned sandgrouse (''Pterocles coronatus'') is a species of bird in the sandgrouse family, the Pteroclidae from North Africa and the Middle East. Description A fairly small sandgrouse which appears rather uniformly coloured from a distance ...
(''Pterocles coronatus'') which has completely dark primaries. The female also has an orange throat region but is generally duller in plumage than the male. The body colour is greyish-brown liberally spotted with small dark markings and with dark patches on the wings, tail and lower belly. The central tail feathers in both sexes are elongated but not to the extent that they are in the
pin-tailed sandgrouse The pin-tailed sandgrouse (''Pterocles alchata'') is a medium large bird in the sandgrouse family. It has a small, pigeon like head and neck and a sturdy, compact body. It has long pointed wings, which are white underneath, a long tail and a fast ...
(''Pterocles alchata''). When flying overhead, a dark belly stripe is visible.


Distribution and habitat

The spotted sandgrouse is found in North Africa and the Middle East. In Africa its range extends through
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
,
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
,
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
,
Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Suda ...
, Sudan,
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Medit ...
, Eritrea,
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
,
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
,
Djibouti Djibouti, ar, جيبوتي ', french: link=no, Djibouti, so, Jabuuti officially the Republic of Djibouti, is a country in the Horn of Africa, bordered by Somalia to the south, Ethiopia to the southwest, Eritrea in the north, and the Red ...
,
Mali Mali (; ), officially the Republic of Mali,, , ff, 𞤈𞤫𞤲𞥆𞤣𞤢𞥄𞤲𞤣𞤭 𞤃𞤢𞥄𞤤𞤭, Renndaandi Maali, italics=no, ar, جمهورية مالي, Jumhūriyyāt Mālī is a landlocked country in West Africa. Mal ...
, Mauritania, Chad and
Niger ) , official_languages = , languages_type = National languagesOman Oman ( ; ar, عُمَان ' ), officially the Sultanate of Oman ( ar, سلْطنةُ عُمان ), is an Arabian country located in southwestern Asia. It is situated on the southeastern coast of the Arabian Peninsula, and spans the mouth of ...
,
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the A ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
,
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
, Syria,
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to the north, Iran to the east, the Persian Gulf and K ...
and
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bordere ...
and its range extends as far as
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's fifth-most populous country, with a population of almost 243 million people, and has the world's second-lar ...
and north west
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
. In 2016, a flock of around a hundred birds arrived at Kutch after a gap of 19 years. It has also been recorded as a vagrant in
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
, the
United Arab Emirates The United Arab Emirates (UAE; ar, اَلْإِمَارَات الْعَرَبِيَة الْمُتَحِدَة ), or simply the Emirates ( ar, الِْإمَارَات ), is a country in Western Asia ( The Middle East). It is located at t ...
and
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula in ...
. It inhabits deserts and semi-arid countryside and is largely resident although there is some local movement of flocks. The population size has not been firmly established but it seems to be stable and the bird seems to be common over most of its extensive range. It is listed as being of "least concern" by the IUCN Red List of Threatened Species.


Behaviour

The spotted sandgrouse has a rapid wing beat and flies swiftly. Its call is a musical "queeto-queeto" which distinguishes it from other species of sandgrouse with similar plumage. The spotted sandgrouse is a ground-dwelling bird and feeds on seeds and other plant material that it finds among the scrubby vegetation of its dry habitat. During the breeding season it is solitary but at other times of year it is gregarious. Flocks move into a new feeding ground after a storm has stimulated new green growth. In the Sahara the spotted sandgrouse are particularly fond of a species of
spurge ''Euphorbia'' is a very large and diverse genus of flowering plants, commonly called spurge, in the family Euphorbiaceae. "Euphorbia" is sometimes used in ordinary English to collectively refer to all members of Euphorbiaceae (in deference to t ...
and concentrate on this until the foliage begins to parch, after which the birds return to their normal diet of seeds. These are abundant on the desert floor, remaining in a dormant state until rain occurs.George, 1978. p. 180 The birds are very wary and easily frightened. Their chief enemy is the
lanner falcon The lanner falcon (''Falco biarmicus'') is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A l ...
which flies rapidly just above the ground and scoops up any unwary bird. The sandgrouse strategy is to have one bird flying high overhead. When it sees an approaching falcon it gives a warning call and the other sandgrouse freeze. So good is their
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
that the
raptor Raptor or RAPTOR may refer to: Animals The word "raptor" refers to several groups of bird-like dinosaurs which primarily capture and subdue/kill prey with their talons. * Raptor (bird) or bird of prey, a bird that primarily hunts and feeds on ...
is unable to detect them and flies on.George, 1978. pp. 157–158 Sandgrouse need to make a daily journey to a drinking hole, which may be many kilometres from their feeding ground. They land a short distance from the water and maintain a sentinel system here too as, beside the falcons, mammal predators may lurk nearby and
nomad A nomad is a member of a community without fixed habitation who regularly moves to and from the same areas. Such groups include hunter-gatherers, pastoral nomads (owning livestock), tinkers and trader nomads. In the twentieth century, the po ...
s may water their herds. When all is safe, another distinctive call from the sentinel bird sends all the others to the pool where their daily water needs are taken up within about fifteen seconds.George, 1978. pp. 159–160 The journey to the waterhole is undertaken at around dawn when the air is cool. Later in the day, when the air temperature may reach over 50 °C, the birds are inactive and have the ability to increase their thermal insulation when the air temperature exceeds their body temperatures. At very high temperatures they also resort to gular fluttering and gaping to cool themselves. At night, several flocks coalesce and fly off into the rocky desert far away from any vegetation. Each bird scrapes itself a shallow sleeping hollow by moving from side to side. There are no
jerboa Jerboas (from ar, جربوع ') are hopping desert rodents found throughout North Africa and Asia, and are members of the family Dipodidae. They tend to live in hot deserts. When chased, jerboas can run at up to . Some species are preyed on b ...
s or other small mammals in these barren wastes so
fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
es and
jackal Jackals are medium-sized canids native to Africa and Eurasia. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed ...
s do not roam there at night and the birds are safe.


Life cycle

In the Sahara, breeding takes place in the hottest part of the summer on a stony desert plain. When choosing a nest site, the female scrapes several trial hollows before selecting one of them. The main criterion for selection seems to be the porous nature of the underlying rock. Spongy rock heats up less in the sun and provides a cool spot to nest. Also desirable are one or two "cover stones" close by, chosen because their dense structure attracts dew at night, moisture which drains into the soil and gets absorbed by the porous rock which helps keep the nest cool by day.George, 1978. pp. 186–188 The nest is made in a shallow depression in the ground without any bedding material and two, occasionally three, eggs are laid. The eggs are elongated ovals in shape, buff with grey and brown blotches and speckles. Their colour and shape makes them difficult to distinguish from the pebbles lying around them. Both parents incubate the eggs and their
cryptic colouration Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
makes them almost invisible when sitting on the nest. The eggs hatch after about 20 days. The young are
precocial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
and already covered in down when they emerge from the eggs. Soon after they are hatched the female leads the chicks to one of the many
wadi Wadi ( ar, وَادِي, wādī), alternatively ''wād'' ( ar, وَاد), North African Arabic Oued, is the Arabic term traditionally referring to a valley. In some instances, it may refer to a wet (ephemeral) riverbed that contains water ...
s that wind across the plains and there she teaches them to peck at and ingest seeds. It is four or five weeks before they are fledged and able to fly. Meanwhile, the problem of supplying them with water is solved by the male which has specially adapted, absorbent down on his belly. While at the waterhole he immerses himself in the water to saturate the plumage which absorbs a quantity that is sufficient for the chicks to last them until the following day. When he leaves the waterhole with his water-laden feathers, the male emits a repeated high-pitched "queet - queet - queet" calls. When the female and chicks pick up the sounds of his approach, they reciprocate, and by this means the male can find his family even when they have moved away from the nest. On his arrival he assumes an upright posture and raises his wings, displaying the wet belly feathers. This is the sign for the chicks to approach and stand underneath him with their beaks upturned and suck the fluid from between the feathers. If danger threatens, the chicks crouch under a plant or any cover that offers, their dappled brown down merging into the desert scene. The parents are very vigilant in defence of their family. If a jackal approaches, an adult gives a warning cry and all crouch down and freeze. Usually the enemy fails to notice them and passes by. If it gets too close, one of the parents tries to lure it away by flapping along the ground, pretending to be injured and helpless. When the jackal has been led far enough away, the adult "recovers" and flies off. After three or four days, the chicks can be left alone while both parents visit the water hole, and the chicks can guide them back with their "queet - queet" cries.George, 1978. pp. 181–182


References


Bibliography

*


External links


Oriental Bird Images: ''Spotted Sandgrouse''
Photos of spotted sandgrouse in the wild. {{Taxonbar, from=Q80832 Pterocles Birds of the Middle East Birds of North Africa Birds described in 1771 Taxonomy articles created by Polbot Taxa named by Carl Linnaeus