Spotted Flycatchers
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The spotted flycatcher (''Muscicapa striata'') is a small passerine bird in the Old World flycatcher family. It breeds in most of Europe and in the
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
to Siberia, and is migratory, wintering in Africa and south western Asia. It is declining in parts of its range. This is an undistinguished looking bird with long wings and tail. The adults have grey-brown upperparts and whitish underparts, with a streaked crown and breast, giving rise to the bird's common name. The legs are short and black, and the bill is black and has the broad but pointed shape typical of aerial insectivores. Juveniles are browner than adults and have spots on the upperparts.


Taxonomy

The spotted flycatcher was described by the German naturalist Peter Simon Pallas in 1764 and given the binomial name ''Motacilla striata''. The genus name ''Muscicapa'' comes from the Latin ''musca'', a fly and ''capere'', to catch. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''striata'' is from the Latin ''striatus'' meaning striated. There are five recognised
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
all of which winter in southern Africa. The breeding range is given below. * ''M. s. striata'' ( Pallas, 1764) – Europe to west Siberia, northwest Africa * ''M. s. inexpectata'' Dementiev, 1932 – Crimea (southern Ukraine) * ''M. s. neumanni'' Poche, 1904 – islands of the Aegean Sea through to the Middle East, the Caucasus, northern Iran and central Siberia * ''M. s. sarudnyi'' Snigirewski, 1928 – eastern Iran and Turkmenistan to the mountains of central Asia and north Pakistan * ''M. s. mongola'' Portenko, 1955 – Mongolia and south-central Siberia Two other subspecies were previously recognised, ''M. s. tyrrhenica'' and ''M. s. balearica''. However, a
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2016 found that they were genetically similar to each other but significantly different from the other spotted flycatcher subspecies. The authors proposed that these insular subspecies should be considered as a separate species. The International Ornithologists' Union has split the species and it is known as the Mediterranean flycatcher, while other taxonomic authorities still consider it to be conspecific.


Description

The spotted flycatcher is a small slim bird, around in length, with a weight of . It has dull grey-brown upperparts and off-white underparts. The crown, throat and breast are streaked with brown while the wings and tail feathers are edged with paler thin margins. The subspecies ''M. s. tyrrhenica'' has paler and warmer plumage on the upperparts, with more diffuse markings on the head and breast. The sexes are alike. Juveniles have ochre-buff spots above and scaly brown spots below.


Behaviour and ecology

Spotted flycatchers hunt from conspicuous perches, making sallies after passing flying insects, and often returning to the same perch. Their upright posture is characteristic. Most passerines moult their primary flight feathers in sequence beginning near the body and proceeding outwards along the wing. The spotted flycatcher is unusual in replacing the outer flight feathers before those nearer the body. The flycatcher's call is a thin, drawn out soft and high pitched ', slightly descending in pitch.


Breeding

They are birds of deciduous woodlands, parks and gardens, with a preference for open areas amongst trees. They build an open nest in a suitable recess, often against a wall, and will readily adapt to an open-fronted nest box. 4-6 eggs are laid. Most European birds cannot discriminate between their own eggs and those of other species. The exception to this are the hosts of the common cuckoo, which have had to evolve this skill as a protection against that nest parasite. The spotted flycatcher shows excellent egg recognition, and it is likely that it was once a host of the cuckoo, but became so good at recognising the intruder's eggs that it ceased to be victimised. A contrast to this is the dunnock, which appears to be a recent cuckoo host, since it does not show any egg discrimination.


Predation

A study conducted at two different locations in southern England found that one third of nests were predated. The
Eurasian jay The Eurasian jay (''Garrulus glandarius'') is a species of passerine bird in the crow family Corvidae. It has pinkish brown plumage with a black stripe on each side of a whitish throat, a bright blue panel on the upper wing and a black tail. The ...
(''Garrulus glandarius'') was the most common aerial predator, consuming both eggs and chicks. The
domestic cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members o ...
(''Felis catus'') predated a small fraction of the nests.


Gallery

File:Muscicapa striata clutch 02.JPG, Clutch File:Grauschnäpper.jpg, Spotted flycatcher nest. File:Juvenile Spotted Flycatcher.JPG, A juvenile flycatcher shortly after leaving the nest. File:Muscicapa striata MWNH 2281.JPG, Egg, Collection Museum Wiesbaden Cuculus canorus canorus MHNT.ZOO.2010.11.150.43.jpg, ''Cuculus canorus canorus'' in a clutch of ''Muscicapa striata'' - MHNT


References


External links


Spotted Flycatcher videos, photos & sounds
on the Internet Bird Collection
Ageing and sexing (PDF; 1.8 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze


* Spotted flycatcher
Species text in The Atlas of Southern African Birds

Xeno-canto: audio recordings of the spotted flycatcher
{{Authority control spotted flycatcher Birds of Europe Birds of Western Asia Birds of Africa spotted flycatcher spotted flycatcher spotted flycatcher spotted flycatcher