''Spiraea virginiana'' is a rare species of flowering plant in the rose family (Rosaceae) known by the common names Virginia meadowsweet and Virginia spiraea. It is native to the southern

Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...

, where it has a distribution scattered across nine states. However, most populations are very small and poor in quality. It is threatened by disturbances in the

hydrology Hydrology () is the scientific study of the movement, distribution, and management of water on Earth and other planets, including the water cycle, water resources, and environmental watershed sustainability. A practitioner of hydrology is calle ...

of its habitat,

introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...

of plants, and other threats. It is a federally listed threatened species of the United States.''Spiraea virginiana''.
NatureServe. Retrieved August 4, 2011. This plant is a

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

growing one to three meters (3–10 feet) tall. It is clonal, with several cloned individuals growing in clumps. The leaves are variable in shape and size and may have smooth or toothed edges. The

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

is a dense

corymb Corymb is a botanical term for an inflorescence with the flowers growing in such a fashion that the outermost are borne on longer pedicels than the inner, bringing all flowers up to a common level. A corymb has a flattish top with a superficial re ...

of many cream-white flowers, each flower about 0.6 centimeters wide. The fruit is a follicle. Flowering occurs in June and July. Though the plant produces many flowers, it rarely reproduces sexually and rarely produces seeds. Instead, it reproduces

vegetatively Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...

by cloning, producing many new sprouts which grow into shrubs.''Spiraea virginiana''.
Center for Plant Conservation. Retrieved August 4, 2011. There are fewer than 30

genotype The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...

s in total, meaning there are fewer than 30 individual sets of genes; all the other plants are clones of one of the 30 types. This plant is a rare member of the flora in the southern Appalachians. It grows in

riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...

habitat, such as

floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...

s and riverbanks. When it was determined to be a threatened species in 1990 there were 24 known occurrences in six states. It could be found at two creeks in

Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...

, the New River,

Little Tennessee River The Little Tennessee River is a tributary of the Tennessee River that flows through the Blue Ridge Mountains from Georgia (U.S. state), Georgia, into North Carolina, and then into Tennessee, in the southeastern United States. It drains portio ...

Nolichucky River The Nolichucky River is a river that flows through Western North Carolina and East Tennessee, in the southeastern United States. Traversing the Pisgah National Forest and the Cherokee National Forest in the Blue Ridge Mountains, the river's wate ...

, and

South Toe River The South Toe River is a river in Yancey County in Western North Carolina. The name Toe is taken from its original name Estatoe, pronounced 'S - ta - toe', a native American name associated with the Estatoe trade route leading down from the NC m ...

North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...

, the Clear Fork and other rivers in

Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...

, the

Russell Fork The Russell Fork, 1975 Board on Geographic Names decision is a U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map accessed June 13, 2011 tributary of the Levisa Fork in southwestern Virginia and sout ...

Pound River The Pound River is a river in the U.S. state of Virginia, running through part of Wise County and through Dickenson County. It runs from the North Fork Pound Reservoir to the John W. Flannagan Dam. Via the Russell Fork, the Levisa Fork, the ...

, New River, and

Guest River The Guest River is a short tributary of the Clinch River in Southwest Virginia, southwestern Virginia in the United States. It is part of the drainage basin, watershed of the Mississippi River by way of the Clinch, Tennessee River, Tennessee and ...

Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...

, the

Bluestone River The Bluestone River is a tributary of the New River, 77 mi (124 km) long, in southwestern Virginia and southern West Virginia in the United States.

Buckhannon River The Buckhannon River is a U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed August 15, 2011 tributary of the Tygart Valley River in north-central West Virginia, USA. Via the Tygart Valley, ...

Gauley River The Gauley River is a river in West Virginia. It merges with the New River to form the Kanawha River, a tributary of the Ohio River. The river features numerous recreational whitewater areas, including those in Gauley River National Recreation A ...

, and

Meadow River The Meadow River is a tributary of the Gauley River, making its headwaters in Greenbrier County and terminating in Nicholas County of West Virginia. It is named for the grassy meadows wetlands which its upper watershed drains, and from which it sp ...

West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...

Rockcastle River The Rockcastle River is a U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map accessed June 8, 2011 river primarily in Rockcastle County, Kentucky, United States. It is a tributary of the Cumberland Ri ...

and other creeks in

Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...

, and other rivers. Besides these populations there are occasional occurrences along rivers and creeks.USFWS
Threatened Status Determined for ''Spiraea virginiana'' (Virginia spiraea).
''Federal Register'' (June 15, 1990). Retrieved August 4, 2011. It is also known from

Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...

and

Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...

.USFWS
''Spiraea virginiana'' Species Profile
Retrieved August 4, 2011. It is thought to have been

extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...

from the state of

Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...

, due to the changes to the

Youghiogheny River The Youghiogheny River , or the Yough (pronounced Yok ) for short, is a U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map accessed August 15, 2011 tributary of the Monongahela River in the U.S. s ...

(the "Yough") caused by

dam A dam is a barrier that stops or restricts the flow of surface water or underground streams. Reservoirs created by dams not only suppress floods but also provide water for activities such as irrigation, human consumption, industrial use, a ...

ming. This shrub grows in cracks in rocks alongside flowing water. It has a fibrous mass of roots and lateral

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

s that anchor it, and even if most of the plant is ripped away during

flood A flood is an overflow of water ( or rarely other fluids) that submerges land that is usually dry. In the sense of "flowing water", the word may also be applied to the inflow of the tide. Floods are an area of study of the discipline hydrol ...

ing, it easily resprouts from remnants of the rhizomes. It is often found growing in

sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...

bedrock In geology, bedrock is solid Rock (geology), rock that lies under loose material (regolith) within the crust (geology), crust of Earth or another terrestrial planet. Definition Bedrock is the solid rock that underlies looser surface mater ...

gravel bar A bar in a river is an elevated region of sediment (such as sand or gravel) that has been deposited by the flow. Types of bars include mid-channel bars (also called braid bars and common in braided rivers), point bars (common in meandering ...

s, nooks between boulders, and in riverborne deposits of debris, rubble, and silt. It is adapted to periodic flooding, scouring,

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is distin ...

, and deposition of litter. Other plants in the habitat may include hazel alder (''Alnus serrulata''),

silky dogwood Silky dogwood is a common name for two species of shrubs, formerly treated as a single species: * ''Cornus amomum'', a more southerly species found in the eastern U.S. * ''Cornus obliqua ''Cornus obliqua'', the blue-fruited dogwood, silky dogwoo ...

(''Cornus amomum''),

jewelweed ''Impatiens'' is a genus of more than 1,000 species of flowering plants, widely distributed throughout the Northern Hemisphere and the tropics. Together with the genus ''Hydrocera'' (one species), ''Impatiens'' make up the family Balsaminaceae. ...

(''Impatiens capensis''), creepers (''Parthenocissus''),

ninebark ''Physocarpus'', commonly called ninebark, is a genus of flowering plants in the family Rosaceae, native to North America (most species) and northeastern Asia (one species). Description ''Physocarpus'' are deciduous shrubs with peeling bark

(''Physocarpus opulifolius''),

sycamore Sycamore is a name which has been applied to several types of trees, but with somewhat similar leaf forms. The name derives from the ancient Greek ' (''sūkomoros'') meaning "fig-mulberry". Species of trees known as sycamore: * ''Acer pseudoplata ...

species (''Platanus''),

blackberries The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus''. The taxonomy of ...

(''Rubus''),

willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...

species (''Salix''), American elderberry (''Sambucus canadensis''),

soapwort ''Saponaria'' is a genus of flowering plants in the family Caryophyllaceae, native to Asia and Europe, and are commonly known as soapworts. They are herbaceous perennials and annuals, some with woody bases. The flowers are abundant, five-petalled ...

s (''Saponaria''), greenbrier species (''Smilax''),

goldenrod Goldenrod is a common name for many species of flowering plants in the sunflower family, Asteraceae, commonly in reference to the genus ''Solidago''. Several genera, such as ''Euthamia'', were formerly included in a broader concept of the genus ...

species (''Solidago''), and poison ivy (''Toxicodendron radicans''). The plant is threatened by its small numbers, alterations in the hydrology of its riparian habitat, and

habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological processes ...

. Populations are isolated from one another, reducing the frequency of its rare sexual reproduction even further. Populations are small; though the plant is adapted to a regime of flooding, unusually large floods have the potential to wipe out small populations. The

invasion An invasion is a military offensive in which large numbers of combatants of one geopolitical entity aggressively enter territory owned by another such entity, generally with the objective of either: conquering; liberating or re-establishing con ...

of introduced plant species, including the shrub's relative, ''Spiraea japonica'' (Japanese spiraea), may be a threat because of

competition Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indivi ...

References

{{Taxonbar, from=Q7577711 virginiana