''Sphex pensylvanicus'', the great black wasp, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of digger wasp. It lives across most of North America and grows to a size of . The larvae feed on living insects that the females paralyze and carry to the underground nest.

Distribution

''S. pensylvanicus'' is distributed across most of the

contiguous United States The contiguous United States (officially the conterminous United States) consists of the 48 adjoining U.S. states and the Federal District of the United States of America. The term excludes the only two non-contiguous states, Alaska and Hawaii ...

and northern

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

. During the late 20th century, its range expanded north to

New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...

and the Canadian provinces of

Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...

and

Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...

Description

''Sphex pensylvanicus'' is a large, black wasp, significantly larger than their congener ''

Sphex ichneumoneus ''Sphex ichneumoneus'', known commonly as the great golden digger wasp or great golden sand digger is a wasp in the family Sphecidae. It is identified by the golden pubescence on its head and thorax, its reddish orange legs, and partly reddish or ...

'' (the great golden digger wasp). Males are smaller than females, at only long compared with typical female sizes of . According to

John Bartram John Bartram (March 23, 1699 – September 22, 1777) was an American botanist, horticulturist, and explorer, based in Philadelphia, Pennsylvania, for most of his career. Swedish botanist and taxonomist Carl Linnaeus said he was the "greatest na ...

, "The sting of this Wasp is painful, but does not swell like others". As well as being larger than ''S. ichneumoneus'', they are also darker, with smoky wings and an entirely black body, where ''S. ichneumoneus'' has yellow wings, red legs, and a partly red abdomen.

Ecology and life cycle

Adult females of ''S. pensylvanicus'' build an underground nest which they provision with various

orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grassho ...

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

s, particularly of the genera ''

Microcentrum ''Microcentrum'' is a genus of phaneropterid bushcrickets, sometimes known as "angle-wing katydids" and found in the Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of Nor ...

'', ''

Amblycorypha ''Amblycorypha'' is a North American genus of round-headed katydids in the family Tettigoniidae Insects in the family Tettigoniidae are commonly called katydids (especially in North America), or bush crickets. They have previously been known ...

'' and ''

Scudderia ''Scudderia'' is a genus of katydids in the subfamily Phaneropterinae. They are sometimes called bush katydids and are 30-38 mm in length. Their range is most of North America, northernmost being in Southern Canada. They are herbivores, with nymp ...

''. Prey are stung three times, once in the neck and twice in the thorax, and are

paralyzed Paralysis (also known as plegia) is a loss of motor function in one or more muscles. Paralysis can also be accompanied by a loss of feeling (sensory loss) in the affected area if there is sensory damage. In the United States, roughly 1 in 50 ...

by the wasp's sting, although they can survive for weeks. The prey are then carried to the nest. While collecting their prey, the females are vulnerable to

kleptoparasitism Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when foo ...

, in which birds, including the

house sparrow The house sparrow (''Passer domesticus'') is a bird of the sparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pale brown and grey, a ...

(''Passer domesticus'') and the

grey catbird The gray catbird (''Dumetella carolinensis''), also spelled grey catbird, is a medium-sized North American and Central American perching bird of the mimid family. It is the only member of the "catbird" genus ''Dumetella''. Like the black catbird ...

(''Dumetella carolinensis''), steal the prey that the wasp has collected. The

eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...

of ''S. pensylvanicus'' are long and wide; they are glued to the underside of the prey insect between the first and second pairs of legs. Each of the several chambers in the nest houses a single

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...

, which consumes 2–6

katydid Insects in the family Tettigoniidae are commonly called katydids (especially in North America), or bush crickets. They have previously been known as "long-horned grasshoppers". More than 8,000 species are known. Part of the suborder Ensifera, t ...

s or

grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...

s. The larval stage lasts 10 days, reaching a pre-pupation size of long by wide. ''S. pensylvanicus'' is an important

pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...

of plants including the

milkweeds ''Asclepias'' is a genus of herbaceous, perennial, flowering plants known as milkweeds, named for their latex, a milky substance containing cardiac glycosides termed cardenolides, exuded where cells are damaged. Most species are toxic to humans ...

Asclepias syriaca ''Asclepias syriaca'', commonly called common milkweed, butterfly flower, silkweed, silky swallow-wort, and Virginia silkweed, is a species of flowering plant. It is native to southern Canada and much of the United States east of the Rocky Moun ...

'' and '' A. incarnata''. It has also been reported on ''

Daucus carota ''Daucus carota'', whose common names include wild carrot, European wild carrot, bird's nest, bishop's lace, and Queen Anne's lace (North America), is a flowering plant in the family Apiaceae. It is native to temperate regions of the Old Wor ...

'', ''

Eryngium yuccifolium ''Eryngium yuccifolium'', known as rattlesnake master, button eryngo, and button snake-root, is a perennial herb of the parsley family native to the tallgrass prairies of central and eastern North America. It grows from Minnesota east to Ohio an ...

,'' ''

Melilotus albus ''Melilotus albus'', known as honey clover, white melilot (UK), Bokhara clover (Australia), white sweetclover (USA), and sweet clover, is a nitrogen-fixing legume in the family Fabaceae. ''Melilotus albus'' is considered a valuable honey plan ...

'', '' Monarda punctata'' , and ''

Pycnanthemum virginianum ''Pycnanthemum virginianum'', the Virginia or common mountain-mint, is a plant in the mint family, Lamiaceae. It is a herbaceous plant with narrow, opposite, simple leaves, on wiry, green stems. The flowers are white with purplish spotting, borne ...

'' ''S. pensylvanicus'' is one of several species of ''

Sphex Wasps of the genus ''Sphex'' (commonly known as digger wasps) are cosmopolitan predators that sting and paralyze prey insects. ''Sphex'' is one of many genera in the old digger wasp family Sphecidae ('' sensu lato''), though most apart from the ...

'' to be parasitized by the

strepsiptera The Strepsiptera are an order of insects with eleven extant families that include about 600 described species. They are endoparasites in other insects, such as bees, wasps, leafhoppers, silverfish, and cockroaches. Females of most species never ...

n '' Paraxenos westwoodi''.

Taxonomic history

''Sphex pensylvanicus'' was the subject of the first article on an insect written by a native of the

New World The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. 3 ...

, when observations made by

on ''S. pensylvanicus'' were presented to the

Royal Society The Royal Society, formally The Royal Society of London for Improving Natural Knowledge, is a learned society and the United Kingdom's national academy of sciences. The society fulfils a number of roles: promoting science and its benefits, re ...

in 1749 by Peter Collinson. ''Sphex pensylvanicus'' was described by

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

in his 1763 work ''

Centuria Insectorum The first page of ''Centuria Insectorum'', as included in ''Amoenitates Academicæ'' ' (Latin, "one hundred insects") is a 1763 taxonomic work by Carl Linnaeus, and defended as a thesis by Boas Johansson; which of the two men should for taxonomi ...

'', using material sent to him by

Charles De Geer Baron Charles de Geer (the family is usually known as De Geer with a capitalized "De" and is pronounced "de yer"); Finspång in Risinge 30 January 1720 – Stockholm 7 March 1778) was a Swedish industrialist and entomologist. Life De Geer, w ...

References

External links

* {{Taxonbar , from=Q7576757 Sphecidae Hymenoptera of North America Insects described in 1763 Taxa named by Carl Linnaeus