Southern Marsh-orchid
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''Dactylorhiza praetermissa'', the southern marsh orchid or leopard marsh orchid, is a commonly occurring
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
an
orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowerin ...
.


Description

''Dactylorhiza praetermissa'' grows to tall, with leaves generally unspotted. The flowers, appearing from May to July, are various shades of pink with variable markings. The basal lip of the flower is rounded. This species is able to form
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
s with other ''Dactylorhiza'' species, and crosses with ''
Dactylorhiza fuchsii ''Dactylorhiza fuchsii'', the common spotted orchid, is a species of flowering plant in the orchid family Orchidaceae. ''Dactylorhiza fuchsii'' is one of Europe's commonest wild orchids. It is widespread across much of Europe, with the range ex ...
'' occur especially often.


Distribution and habitat

It is native to northern and central
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
(
Britain Britain most often refers to: * The United Kingdom, a sovereign state in Europe comprising the island of Great Britain, the north-eastern part of the island of Ireland and many smaller islands * Great Britain, the largest island in the United King ...
,
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of Denmark , establish ...
,
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and the ...
, Sweden,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
,
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan area ...
,
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...
and Latvia). It is also reportedly naturalized in
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
and in parts of
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
(
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central C ...
and Newfoundland).Kew World Checklist of Selected Plant Families
/ref> This species is found close to water, in damp alkaline meadows, by ponds, lakes or reservoirs and in dune slacks.


Ecology

The flowers of this species are pollinated by insects including the
cuckoo bee The term cuckoo bee is used for a variety of different bee lineages which have evolved the kleptoparasitic behaviour of laying their eggs in the nests of other bees, reminiscent of the behavior of cuckoo birds. The name is perhaps best applied to ...
and
skipper butterfly Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...
. ''Dactylorhiza'' are known to be
mycorrhizal   A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the pla ...
generalists. ''D. praetermissa'' has been shown to benefit from association with fungal species in the genus ''
Rhizoctonia ''Rhizoctonia'' is a genus of fungi in the order Cantharellales. Species form thin, effused, corticioid basidiocarps (fruit bodies), but are most frequently found in their sterile, anamorphic state. ''Rhizoctonia'' species are saprotrophic, bu ...
'' and others in the ''
Tulasnellaceae The Tulasnellaceae are a family of fungi in the order Cantharellales. The family comprises mainly effused (patch-forming) fungi formerly referred to the " jelly fungi" or heterobasidiomycetes. Species are wood- or litter-rotting saprotrophs, but ...
'' family.PLoS One - Variation in Mycorrhizal Associations with Tulasnelloid Fungi Among Populations of Five Dactylorhiza Species
/ref>


Subspecies and varieties

Many names for infraspecific taxa have been proposed. At of June 2014, the following are recognized: #''Dactylorhiza praetermissa'' var. ''junialis'' (Verm.) Senghas - Britain, France, Belgium, Netherlands, Canada #''Dactylorhiza praetermissa'' subsp. ''osiliensis'' (Pikner) Kreutz - Sweden, Estonia #''Dactylorhiza praetermissa'' subsp. ''praetermissa'' - Britain, Denmark, Norway, Netherlands, Belgium, Germany, France, Italy #''Dactylorhiza praetermissa'' subsp. ''schoenophila'' R.M.Bateman & Denholm - Britain


References


External links


Biohorizons - An investigation to determine variation in marsh orchid (Dactylorhiza) populations at Moses Gate Country Park, BoltonFirst Nature - Dactylorhiza praetermissa
praetermissa Orchids of Europe Plants described in 1914 {{orchidoideae-stub