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The southern grasshopper mouse or scorpion mouse (''Onychomys torridus'') is a species of predatory rodent in the family
Cricetidae The Cricetidae are a family of rodents in the large and complex superfamily Muroidea. It includes true hamsters, voles, lemmings, muskrats, and New World rats and mice. At almost 608 species, it is the second-largest family of mammals, and has m ...
, native to
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
and the states of
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
,
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
,
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
, and
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
. Notable for its resistance to venom, it routinely preys on the highly venomous
Arizona bark scorpion :''The striped bark scorpion and the closely related Baja California bark scorpion are also called bark scorpions.'' The Arizona bark scorpion (''Centruroides sculpturatus'', once included in ''Centruroides exilicauda'') is a small light brown s ...
.


Description

The southern grasshopper mouse is a robust, small, nocturnal species that typically forms monogamous pairs. They have a short tail, growing to a total length of . The head, back and sides are pinkish- or grayish-brown while the underparts are white, the two colors being distinctly separated. The tail is club-shaped, short and broad, the anterior part being the same color as the body and the tip being white.


Biology

The southern grasshopper mouse feeds almost entirely on
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s, such as
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s,
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...
s and
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always end ...
s. In addition to arthropods, it is known to prey upon the
little pocket mouse The little pocket mouse (''Perognathus longimembris'') is a species of rodent in the family Heteromyidae. It is found in Baja California and Sonora in Mexico and in Arizona, California, Idaho, Nevada, Oregon and Utah in the United States. Its nat ...
(''Perognathus longimembris'') and the
western harvest mouse The western harvest mouse (''Reithrodontomys megalotis'') is a small neotomine mouse native to most of the western United States. Many authorities consider the endangered salt marsh harvest mouse to be a subspecies, but the two are now usually ...
(''Reithrodontomys megalotis''). In the arid regions inhabited by the southern grasshopper mouse, the Arizona bark scorpion is plentiful and avoided by most predators because of the very painful sting it can inflict. The grasshopper mouse can feed on this scorpion with impunity even when the scorpion stings it repeatedly in the face. Researchers have found that a neural mechanism is involved that blocks the sending of pain messages to the brain. This mouse can normally feel pain from other sources, but can be temporarily insensitive to these stimuli after a dose of venom from a bark scorpion. This pain modulation is activated soon after the scorpion sting; the mouse does briefly feel pain from the sting, but the scorpion venom binds to the transmembrane channel otNav1.8. Specifically, the venom binds to a glutamic acid residue located within the channel protein, thus preventing neuron action potentials from firing and acting as a type of temporary anesthetic and pain reliever for the mouse. When this glutamic acid was replaced with hydrophilic glutamine residue, the venom did not bind with the protein, and the pain modulation ability was lost. Females are able to produce several litters in a year, but their reproductive life does not usually last into their second year. The average litter size is 2.6 (range 1 to 5), produced after an average gestation period of 29 days. The young are born naked, with pink translucent skin. One day later the pigment is developing on the dorsal surface and by day ten, hairs long are present on the back. The eyes open on day 15 and solid food is being consumed by day 19. The females give birth to their first litter when four or five months old.


Ecology

Southern grasshopper mice are unique among other North American rodents in the family Cricetidae in that their diet consists mostly of arthropods. They are usually found in areas with lower population densities. Several studies have demonstrated that the reproductive attributes of this species allows them to be naturally maintained at lower population densities. Females tend to be sexually active only during one breeding season, which is due to their rapid reproductive aging following their first year. Moreover, males' unique pauses in their testicular activity during the breeding season might also contribute to the lower population densities in which they live. The natural habitat of southern grasshopper mice include hot, arid valleys, and scrub deserts. A male and female along with their offspring occupy a larger home range for burrowing compared to males and females alone. On average, males have a home range of 7.8 acres, while females occupy an area of 5.9 acres. They tend to maintain a specific place of residency for a longer period of time compared to other mammals. The home ranges between southern grasshopper mice are not too far from each other, with an average maximum distance of 118.4± 8.0 meters. However, adults males tend to be very territorial and their efficient spacing is facilitated by vocalizations they make at night. In a study done by Denise H. Frank and Edward J. Heske, the home range of southern grasshopper mice in the Chihuahuan Desert were observed.Frank, D. H., & Heske, E. J. (1992). Seasonal changes in space use patterns in the southern grasshopper mouse, Onychomys torridus torridus. ''Journal of Mammalogy'', ''73''(2), 292-298. The home range of males during the non-breeding season were found to be smaller than those held by males during the breeding season. In contrast, there was not a significant difference between the home ranges of females during the breeding and non-breeding seasons. Male ranges during the reproductive season were some two to three times larger than those of adult females during the reproductive season. There are no currently known conservation issues associated with southern grasshopper mice, largely due to its wide distribution and tolerance for disturbances caused by arthropods.Guevara-Carrizales, A. A., Escobar-Flores, J., & Martínez-Gallardo, R. (2012). Marginal record of the southern grasshopper mouse Onychomys torridus in Baja California, México. ''Western North American Naturalist'', ''72''(3), 416-417. However, some disturbances have been observed in their low density populations. Many more population studies will need to be performed in order to gain further insight into the conservation status of this species.


Predatory behavior

Southern grasshopper mice are a predatory, and very aggressive rodent species.McCarty, R., & Southwick, C. H. (1981). Food deprivation: Effects on the predatory behavior of southern grasshopper mice (Onychomys torridus).''Aggressive Behavior'', ''7''(2), 123-130. In a study done by Richard McCarty and Charles H. Southwick, these grasshopper mice were deprived of food and their predatory behavior against laboratory mice and crickets was observed. After 48 hours of food deprivation, southern grasshopper mice spent more time eating crickets than the laboratory food they were provided. Live crickets were preferred to deceased specimens, which might suggest that prey movement might be a catalyst and cue for predatory attack. There was no gender related difference in how much time the mice spent feeding on crickets. Highly aggressive behaviors were displayed towards laboratory mice in the same study, such as biting the back and tail of their prey. After two days of being around the laboratory mice, six of the 43 male grasshopper mice and 22 of the 45 females grasshopper mice killed and ate certain parts of the laboratory mice. This difference in predatory behavior between the male and female grasshopper mice was significant. In their study, both McCarty and Southwick concluded that food deprivation and the type of prey affected the intensity and aggressiveness of the predatory attacks observed by southern grasshopper mice.


Brain and body size relation

Relative brain size tends to vary with food preferences.Mann, M. D., & Towe, A. L. (2003). Brain-body size relations in grasshopper mice. ''Brain, behavior and evolution'', ''62''(1), 13-18. Brain size is usually smallest in folivores, larger in both granivores and insectivores, and largest in generalists. Measurements of cranial volume of museum specimens of southern grasshopper mouse have shown that on average, females were heavier and longer than males, but their cranial volume was about 3% less than that of males. However, this was not a significant difference. This pattern of brain-body relationship can also be seen in northern grasshopper mice. Moreover, the tail length of southern grasshopper mice made up an average of 33.9% of its body length. Their tail length measures an average of 4.58 ± 0.36 cm in females, and 4.50 ± 0.39 cm in males. Average tail length does not vary greatly between the sexes. On average, female southern grasshopper mice are able to deal with a larger body size without increasing their brain size, when compared to males. There is still not enough evidence to explain the difference in body size and brain between males and females. It is possible that there are different demands on both sexes, which might contribute to this difference. These studies were also only done in the laboratory, and there might be sex-specific stresses in their natural habitat that influence brain growth. However, sexual dimorphism is quite common and has been observed in the central nervous system of other mammals.


References

{{Taxonbar, from=Q277147 Onychomys Mammals described in 1874 Taxonomy articles created by Polbot