Soricomorpha (from

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

"shrew-form") is a formerly used

taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

within the

class Class or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used differentl ...

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s. In the past it formed a significant group within the former order

Insectivora The order Insectivora (from Latin ''insectum'' "insect" and ''vorare'' "to eat") is a now-abandoned biological grouping within the class of mammals. Some species have now been moved out, leaving the remaining ones in the order Eulipotyphla, w ...

. However, Insectivora was shown to be

polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...

and various new orders were split off from it, including

Afrosoricida The order Afrosoricida (a Latin-Greek compound name which means "looking like African shrews") contains the golden moles of Southern Africa, the otter shrews of equatorial Africa and the tenrecs of Madagascar. These three families of small mamma ...

(

tenrecs A tenrec is any species of mammal within the afrotherian family Tenrecidae endemic to Madagascar. Tenrecs are wildly diverse; as a result of convergent evolution some resemble hedgehogs, shrews, opossums, rats, and mice. They occupy aquatic ...

golden mole Golden moles are small insectivorous burrowing mammals endemic to Sub-Saharan Africa. They comprise the family Chrysochloridae and as such they are taxonomically distinct from the true moles, family Talpidae, and other mole-like families, all o ...

s, otter shrews), Macroscelidea (elephant shrews), and

Erinaceomorpha Erinaceidae is a family in the order Eulipotyphla, consisting of the hedgehogs and moonrats. Until recently, it was assigned to the order Erinaceomorpha, which has been subsumed with the paraphyletic Soricomorpha into Eulipotyphla. Eulipotyph ...

(

hedgehog A hedgehog is a spiny mammal of the subfamily Erinaceinae, in the eulipotyphlan family Erinaceidae. There are seventeen species of hedgehog in five genera found throughout parts of Europe, Asia, and Africa, and in New Zealand by introducti ...

s and

gymnure Gymnures, also called hairy hedgehogs or moonrats, are mammals belonging to the subfamily Galericinae, in the family Erinaceidae and the order Eulipotyphla. Gymnures resemble rats but are not closely related as they are not rodents; they are ...

s), with the four remaining extant and recent

families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...

of Soricomorpha shown here then being treated as a separate order. Insectivora was left empty and disbanded. Subsequently, Soricomorpha itself was shown to be

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

, because Soricidae shared a more recent common ancestor with Erinaceidae than with other soricomorphs. The combination of Soricomorpha and Erinaceidae, referred to as order

Eulipotyphla Eulipotyphla (, which means "truly fat and blind") is an order of mammals suggested by molecular methods of phylogenetic reconstruction, which includes the laurasiatherian members of the now-invalid polyphyletic order Lipotyphla, but not the afr ...

, has been shown to be monophyletic. Living members of the group range in size from the

Etruscan shrew The Etruscan shrew (''Suncus etruscus''), also known as the Etruscan pygmy shrew or the white-toothed pygmy shrew, is the smallest known extant mammal by mass, weighing only about on average. (The bumblebee bat is regarded as the smallest mammal ...

, at about 3.5 cm and 2 grams, to the

Cuban solenodon The Cuban solenodon or ''almiquí'' (''Atopogale cubana'') is a small, furry, shrew-like mammal endemic to mountainous forests on Cuba. It is the only species in the genus ''Atopogale''. An elusive animal, it lives in burrows and is only active ...

, at about 32 cm and 1 kg. * Soricomorpha ** Family Soricidae (shrews) *** Subfamily

Crocidurinae The white-toothed shrews or Crocidurinae are one of three subfamilies of the shrew family Soricidae. The outer layer of these shrews' teeth is white, unlike that of the red-toothed shrews. These species are typically found in Africa and souther ...

: (white-toothed shrews) *** Subfamily

Soricinae The red-toothed shrews of the subfamily Soricinae are one of three living subfamilies of shrews, along with Crocidurinae (white-toothed shrews) and Myosoricinae (African white-toothed shrews). In addition, the family contains the extinct subfamil ...

: (red-toothed shrews) *** Subfamily

Myosoricinae According to the current taxonomy, the Myosoricinae are a subfamily of shrews. As such, they form one of three main types of shrews, the other two being the red-toothed shrews and the white-toothed shrews. They are the only one of the three to ...

: (African white-toothed shrews) ** Family

Talpidae The family Talpidae () includes the moles (some of whom are called shrew moles and desmans) who are small insectivorous mammals of the order Eulipotyphla. Talpids are all digging animals to various degrees: moles are completely subterranean a ...

: (

moles Moles can refer to: * Moles de Xert, a mountain range in the Baix Maestrat comarca, Valencian Community, Spain * The Moles (Australian band) *The Moles, alter ego of Scottish band Simon Dupree and the Big Sound People *Abraham Moles, French engin ...

and close relatives) *** Subfamily

Scalopinae The Scalopinae, or New World moles, are one of three subfamilies of the family Talpidae, which consists of moles and mole-like animals; the other two subfamilies being the Old World talpids (the Talpinae) and the Chinese shrew-like moles (Uropsil ...

(New World moles and close relatives) *** Subfamily

Talpinae The subfamily Talpinae, sometimes called "Old World moles" or "Old World moles and relatives", is one of three subfamilies of the mole family Talpidae, the others being the Scalopinae, or New World moles, and the Uropsilinae, or shrew-like moles. ...

(Old World moles and close relatives) *** Subfamily

Uropsilinae The shrew moles or shrew-like moles (''Uropsilus'') are shrew-like members of Talpidae, the mole family of mammals endemic to the forested, high-alpine region bordering China, Myanmar, and Vietnam. They possess a long snout, a long slender tail, ...

(Chinese shrew-like moles) ** Family

Solenodontidae Solenodons (from el, τέλειος , 'channel' or 'pipe' and el, ὀδούς , 'tooth') are venomous, nocturnal, burrowing, insectivorous mammals belonging to the family Solenodontidae . The two living solenodon species are the Cuban solen ...

: solenodons (rare primitive eulipotyphlans of the Caribbean; two extant species) ** Family †

Nesophontidae ''Nesophontes'', sometimes called West Indies shrews, is the sole genus of the extinct, monotypic mammal family Nesophontidae in the order Eulipotyphla. These animals were small insectivores, about 5 to 15 cm long, with a long slender snout ...

: West Indian shrews (recently extinct eulipotyphlans of the Caribbean) ** Family † Heterosoricidae *** genus † Atasorex *** genus †

Dinosorex ''Dinosorex'' is an extinct eulipotyphlan genus, popularly referred to as giant terror shrews due to their fearsome lower incisors. ''Dinosorex'' lived in Europe from the late Oligocene or early Miocene to the late Miocene, with a range that stre ...

*** genus †'' Domnina'' *** genus † Gobisorex *** genus † Heterosorex *** genus † Ingentisorex *** genus †

Lusorex ''Lusorex'' is an extinct genus of heterosoricid shrew that lived in China during the Miocene. The only species is ''L. taishanensis'', fossils of which are kept at the Paleozoological Museum of China The Paleozoological Museum of China (PMC; ) ...

*** genus † Paradomnina *** genus † Quercysorex ** Family †

Nyctitheriidae Nyctitheriidae is a family of extinct eulipotyphlan insectivores known from the Paleocene and Eocene epochs of North America and Asia and persisting into the Oligocene of Europe. Several genera, including '' Nyctitherium'', '' Paradoxonycteris' ...

References

{{Taxonbar, from=Q834919 Taxa named by William King Gregory Extant Eocene first appearances Paraphyletic groups Eulipotyphla Obsolete mammal taxa