The sooty-headed wren (''Pheugopedius spadix'') is a species of

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

in the family Troglodytidae. It is found in

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...

and

Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...

Taxonomy and systematics

The sooty-headed wren was previously treated as

conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...

with the black-throated wren (''Pheugopedius atrogularis'') but significant differences in their plumage, song, and ranges demanded their separation. According to the

International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...

(IOC) the sooty-headed wren is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

. However, the

Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...

BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...

(BLI), and the

Cornell Lab of Ornithology The Cornell Lab of Ornithology is a member-supported unit of Cornell University in Ithaca, New York, which studies birds and other wildlife. It is housed in the Imogene Powers Johnson Center for Birds and Biodiversity in Sapsucker Woods Sanctuar ...

's ''Birds of the World'' recognize two subspecies, the nominate ''Pheugopedius spadix spadix'' and ''P. s. xerampelinus''.Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, B. L. Sullivan, and C. L. Wood. 2019. The eBird/Clements Checklist of Birds of the World: v2019. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 15, 2019HBW and BirdLife International (2020) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021Kroodsma, D. E. and D. Brewer (2020). Sooty-headed Wren (''Pheugopedius spadix''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.sohwre1.01 retrieved June 3, 2021

Description

The sooty-headed wren is long. The nominate adults' crown, shoulders, and back are dull blackish and the rump bright chestnut. Their tail is chestnut with bold black bars. Much of their face is black with white streaks. Their chin and throat are black and contrast with the bright chestnut lower throat and chest. The belly is a duller grayish brown between reddish brown flanks. The juvenile is duller than the adult and has less contrast between the areas of different color. ''P. s. xerampelinus'' is overall paler than the nominate and has less extensive reddish brown on the flanks.

Distribution and habitat

The nominate sooty-headed wren is found in Colombia, in the Pacific lowlands from southern

Chocó Department Choco Department is a department of Western Colombia known for its large Afro-Colombian population. It is in the west of the country, and is the only Colombian department to have coastlines on both the Pacific Ocean and the Atlantic Ocean. It ...

to Nariño Department and east from there into the central

Magdalena Valley The Magdalena River Valley ( es, Valle del Río Magdalena) is a valley in Colombia located within the Colombian Andes. The valley is specifically situated between the Central and Eastern Ranges and crossed by the river of the same name, the Magdale ...

and some isolated mountain ranges. ''P. s. xerampelinus'' is found on the Pacific slope of far southern Panama's

Darién Province Darién (, , ) is a province in Panama whose capital city is La Palma. With an area of , it is located at the eastern end of the country and bordered to the north by the province of Panamá and the region of Kuna Yala. To the south, it is border ...

. The species inhabits the interior and edges of humid forest,

cloudforest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud c ...

, and dense

secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...

. It favors areas with heavy moss growth. In elevation it mostly ranges between but can be found as low as in Colombia.

Behavior

Feeding

The sooty-headed wren usually forages in pairs, staying low in the vegetation, and seldom joins

mixed-species foraging flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...

s. It has been observed following

army ant The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limit ...

swarms. Its prey is mostly or entirely invertebrates including spiders and adult and larval insects.

Breeding

Though little information has been published about the sooty-headed wren's breeding

phenology Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonality, seasonal and interannual variations in climate, as well as environmental factor, habitat factors (such as elevation). Examples includ ...

, it appears to have a breeding season spanning from March to December. Its nest is a ball made of coarse material and leaves with a side entrance.

Vocalization

Both sexes of the sooty-headed wren sing, "a series of...gurgling whistles

Status

The

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

has assessed the sooty-headed wren as being of Least Concern. It is " irly common in suitable habitat in Colombia."

References

{{Taxonbar, from=Q4749241 Pheugopedius Birds of Colombia Birds of the Tumbes-Chocó-Magdalena Birds described in 1910 Taxonomy articles created by Polbot