Somosaguas Paleontological Site
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The Somosaguas fossil site is located in the Community of Madrid, in the Somosaguas campus, within the municipality of Pozuelo de Alarcón ( Spain). The fossils found belong to the fauna of the Aragonian continental stage (middle Miocene), about 14 million years ago. The sediments where the remains are found were deposited in an alluvial fan sedimentary environment. The site itself is formed by two zones with fossil presence, separated by : Somosaguas Norte, which occupies an upper stratigraphic position, and Somosaguas Sur. The site was discovered in 1989 by a geology student near the Faculty of Political Sciences of the Complutense University, when he observed bone fragments in the ground. The student reported the discovery to Professor Nieves López Martínez, who verified that the remains were fossils, and in turn reported it to the Department of Paleobiology of the Museo Nacional de Ciencias Naturales.


History

In 1989 a geology student at the Complutense University of Madrid, Francisco Hernández Arteaga, was waiting for his girlfriend next to the Faculty of Political Science at the Somosaguas campus when he discovered skeletal remains on the ground. He did not report the discovery until 1996. Once the discovery was reported, members of the Paleobiology Department of the National Museum of Natural Sciences carried out prospections in the area, the results of which were included in a report presented to the
Ministry of Culture of the Community of Madrid Ministry may refer to: Government * Ministry (collective executive), the complete body of government ministers under the leadership of a prime minister * Ministry (government department), a department of a government Religion * Christian m ...
, and the site was included in the
paleontological chart Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fo ...
. The first paleontological excavation took place in 1998, in which Professor Nieves López led a team of students from the Faculty of Geological Sciences of Madrid, with the financial support of the Rector's Office of the Complutense University, and with the collaboration of the National Museum of Natural Sciences. Since then, excavations have been carried out regularly by students and since 2000 a fortnightly work period has been established in April or May, in addition to open days, workshops and exhibitions.


Geology

The site is located on Miocene materials that were deposited by a system of alluvial fans, which were formed due to the erosion suffered by the granitic and metamorphic rocks of the
Central System The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...
. These sediments outcrop from the south of the urban area of Madrid to the contact with the plutonic and metamorphic materials of the sierra. The sediments present in the deposit have a potency (sediment thickness) of 6 m and consist of
arkose Arkose () or arkosic sandstone is a detrital sedimentary rock, specifically a type of sandstone containing at least 25% feldspar. Arkosic sand is sand that is similarly rich in feldspar, and thus the potential precursor of arkose. Quartz is c ...
s with clayey
matrix Matrix most commonly refers to: * ''The Matrix'' (franchise), an American media franchise ** '' The Matrix'', a 1999 science-fiction action film ** "The Matrix", a fictional setting, a virtual reality environment, within ''The Matrix'' (franchi ...
and rock fragments with intercalations of brown clays and
mica Micas ( ) are a group of silicate minerals whose outstanding physical characteristic is that individual mica crystals can easily be split into extremely thin elastic plates. This characteristic is described as perfect basal cleavage. Mica is ...
ceous sands, distinguishing three sections. * Section 1 (T1): it forms the wall of the series and is thick. It is formed by arkoses with a clayey matrix, and in this section of the Somosaguas Sur deposit there are a large number of micro and macrovertebrate fossils. * Section 2 (T2): alternating levels of micaceous sands and brown clays, with a maximum thickness of . * Section 3 (T3): At the top of the series, it has a thickness of more than , formed by arkoses with clayey matrix. The macrovertebrate fossils of Somosaguas Norte are found in this section.


Fossils

The Somosaguas Sur site has a great richness of micromammal fossils (more than 400 fossil teeth were found in of sediment) that have allowed us to assign it to the Middle Aragonian E biozone (13.75–14.10 million years ago).A biozone is a set of sedimentary rocks characterized by the presence of fossils of a certain taxon or taxa, which lived for a time shorter than the duration of a geologic stage. () The taxa identified belong to different groups: Gliridae ('' Microdyromys koenigswaldi, Microdyromys monspeliensis'' and '' Armantomys tricristatus''), Sciuridae ('' Heteroxerus grivensis'' and '' Heteroxerus rubricati''), Cricetidae ('' Megacricetodon collongensis'', '' Democricetodon darocensis'', '' Cricetodon soriae'' and ''Democricetodon'' ''lacombai''), Insectivora ( ''Miosorex'' ''grivensis'' and ''
Galerix exilis ''Galerix'' is a prehistoric genus of gymnures. Fossils of these hedgehog-like creatures are found in Europe, Africa, and Asia. Taxonomy Nine species are recognized as members of the genus: *''Galerix aurelianensis'' *'' Galerix exilis'' *'' ...
''), Lagomorpha ('' Lagopsis penai'' and ''Prolagus'' ''oeningensis'') and Reptilia ('' Chelonia'', ''
Lacertidae The Lacertidae are the family (biology), family of the wall lizards, true lizards, or sometimes simply lacertas, which are native to Afro-Eurasia. It is a diverse family with at least 300 species in 39 genera. They represent the dominant group o ...
'' and '' Anguidae''). Among the macrovertebrates, remains of the mastodon '' Gomphotherium angustidens'' have been recovered; from the family Rhinocerotidae, the taxon ''Alicornops'' ''simorrense'' has been identified; from the family Equidae, remains of '' Anchitherium'' have been found; and among the ruminants, there are fossils of the
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
''
Heteroprox ''Heteroprox'' is an extinct genus of deer from the Miocene of Europe. Description ''Heteroprox'' was a mid-sized deer that would have been similar in appearance to a muntjac Muntjacs ( ), also known as the barking deer or rib-faced deer, ( ...
'' ( Cervidae), '' Micromeryx'' ( Moschidae) and '' Tethytragus'' ( Bovidae). There is also a
suid The Unix access rights flags setuid and setgid (short for ''set user identity'' and ''set group identity'') allow users to run an executable with the file system permissions of the executable's owner or group respectively and to change behaviour ...
, ''
Conohyus ''Conohyus'' was an extinct genus of suid that existed during the Miocene in Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and tr ...
''. The fossils present in Somosaguas Norte correspond mostly to macrovertebrates, although microvertebrates have also been found, such as the lagomorphs '' Lagopsis'' and ''
Prolagus oeningensis ''Prolagus oeningensis'' is an extinct lagomorph and the type species of its genus, ''Prolagus''. It lived from 15.97 to 7.75 Ma, existing for about 8 million years. Range The species has been found in various locations in Europe and Asia. It wa ...
'' and the
cricetid The Cricetidae are a family (biology), family of rodents in the large and complex superfamily Muroidea. It includes true hamsters, voles, lemmings, Muskrat, muskrats, and New World rats and mice. At almost 608 species, it is the second-largest Fa ...
'' Democricetodoninae''. As for macrovertebrates, remains of the mastodon '' Gomphotherium angustidens'' have been documented; fossils of the carnivores ''
Hemicyon ''Hemicyon'', also known as the "dog-bear" (literally "half dog", from Greek (half) + (dog)), is an extinct genus of hemicyonine bear, which probably originated in Eurasia but was found in Europe, Asia and North America during the Miocen ...
'', ''
Amphicyon ''Amphicyon'' ("ambiguous dog") is an extinct genus of large carnivorous bone-crushing mammals, popularly known as bear dogs, of the family Amphicyonidae, subfamily Amphicyoninae, from the Burdigalian Epoch until the late Pliocene, with the creat ...
'', '' Pseudaelurus'' and an undetermined mustelid; the equid '' Anchitherium''; the rhinoceros '' Prosantorhinus douvillei''; the ruminants ''
Heteroprox ''Heteroprox'' is an extinct genus of deer from the Miocene of Europe. Description ''Heteroprox'' was a mid-sized deer that would have been similar in appearance to a muntjac Muntjacs ( ), also known as the barking deer or rib-faced deer, ( ...
'', '' Tethytragus'' and '' Micromeryx'' and the suid '' Conohyus simorrensis''. Bird and chelonian remains have also been recovered.


Taphonomy, paleoclimatology and paleoecology

Based on the taphonomic study of the Somosaguas Norte site, the following observations have been made: there are a large number of splinters (small bone fragments) from two different types of bone: cancellous bone splinters and compact bone, and they seem to indicate that the bone remains have undergone intense weathering processes. The presence of some fragile remains and intact surfaces has also been documented, as well as the presence of very altered remains that form powdery masses. In addition, signs of abrasion and mineral replacement have been observed in the remains. It seems that the main processes that have occurred in the deposit from the taphonomic point of view are: * Presence of different thanatoceonosis with different degrees of preservation. * Concentration of the remains. * Compaction of the sediment during fossildiagenesis. *
Mineralization Mineralization may refer to: * Mineralization (biology), when an inorganic substance precipitates in an organic matrix ** Biomineralization, a form of mineralization ** Mineralization of bone, an example of mineralization ** Mineralized tissues are ...
of the bones. * Cementation, cavity filling,
decalcification Bone decalcification is the softening of bones due to the removal of calcium ions, and can be performed as a histological technique to study bones and extract DNA. This process also occurs naturally during bone development and growth, and when uninh ...
and decomposition of bones. * Differential compaction causing fractures in certain remains. Isotopic analysis of the fossil remains has shown that a cooling of the climate and an increase in aridity were taking place at that time. Calculations indicate a temperature drop of , coinciding with the drop in temperatures recorded during the Miocene as a consequence of the re-establishment of the polar ice cap in Antarctica 14 million years ago. Also the presence of dioctahedral smectite in the sediments indicates the presence of a very long dry season. By studying the Ba/ Ca ratio in tooth enamel samples of different taxa, it has been concluded that the equids of the genus ''Anchitherium'' ingested a higher proportion of herbaceous plants than the mastodons of the genus ''Gomphotherium'' and that the feeding habits of the suid ''Conohyus simorrensis'' were more omnivorous. Among the fossil remains found, the abundance of the different taxa has been calculated from the bone remains identified between 1998 and 2006, with 47.47% of macrofaunal remains and 52.53% of microfaunal remains in the site. Within the macrofauna, ruminants are represented with 14.4%, the genus ''Amchitherium'' with 12.7%, '' Gomphotherium angustidens'' with 11.1%, carnivores with 3% as well as '' Conohyus simorrensis'' and finally '' Prosantorhinus douvillei'' represents 2.6% of the taxa present. The most abundant taxon is the microvertebrate '' Megacricetodon collongensis'' (24%). As for other microvertebrates, there are '' Democricetodon larteti'' (11.2%), '' Democricetodon'' (0.1%), '' Cricetodon soriae'' (0.3%), '' Armantomys tricristatus'' (5%), '' Microdyromys koenigswaldi'' and '' M. monspeliensis'' (4.2%), '' Heteroxerus grivensis'' (4.8%), ''
Prolagus oeningensis ''Prolagus oeningensis'' is an extinct lagomorph and the type species of its genus, ''Prolagus''. It lived from 15.97 to 7.75 Ma, existing for about 8 million years. Range The species has been found in various locations in Europe and Asia. It wa ...
'' (0.1%), '' Lagopsis penai'' (1.5%), ''
Galerix exilis ''Galerix'' is a prehistoric genus of gymnures. Fossils of these hedgehog-like creatures are found in Europe, Africa, and Asia. Taxonomy Nine species are recognized as members of the genus: *''Galerix aurelianensis'' *'' Galerix exilis'' *'' ...
'' (1.5%), '' Miosorex grivensis'' (0.6%) and non-mammals (0.7%).


See also

* List of fossil sites


Notes


References


External links

{{Commons category, Somosaguas fossil site Pozuelo de Alarcón Miocene Paleontological sites