Somatochlora
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''Somatochlora'', or the striped emeralds, is a genus of
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threa ...
in the family
Corduliidae The Corduliidae, also knowns as the emeralds, emerald dragonflies or green-eyed skimmers, is a family of dragonflies. These dragonflies are usually black or dark brown with areas of metallic green or yellow, and most of them have large, emerald-gr ...
with 42 described species found across the Northern Hemisphere.


Taxonomy

The name ''Somatochlora'' is derived from the Greek ''soma'' (body) and ''khloros'' (green). The species '' Corduliochlora borisi'' was formerly treated as a member of ''Somatochlora''.


Description

Members of this genus are medium-sized dragonflies with dark bodies and a metallic green lustre. The eyes are brilliant green, and many species have dull to bright yellow markings on the
thorax The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the cre ...
and/or
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
. The abdomens of males are distinctive, with the first two segments bulbous-shaped, the third constricted, and the rest of the abdomen club-shaped with a straight ending. Females have abdomens with straighter sides. Identifying these dragonflies to species can be difficult. The cerci of males, on the tip of the abdomen, are distinctively shaped in each species, as are the subgenital plates on female abdomens. In some species, the subgenital plate is large and projecting, and is used as a "pseudo-
ovipositor The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
" for inserting eggs into a substrate.


Distribution

''Somatochlora'' are found across the Northern Hemisphere in Europe, Asia and North America, with some species extending into arctic regions north of the
treeline The tree line is the edge of the habitat at which trees are capable of growing. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions (usually cold temperatures, extreme snowp ...
. Some species extend south to
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
, northern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
, northern
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
and the southern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
. At least one species, ''S. semicircularis, may be found at altitudes of up to 3700 metres. In North America, most species live in the
boreal forest Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, ...
and/or
Appalachian mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...
.


Life History

''Somatochlora'' larvae typically live in
bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and muskeg; a ...
s,
fen A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetlands along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires. T ...
s and/or forest
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...
s, with some species found in
lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...
s. They do not occur in marshy
pond A pond is an area filled with water, either natural or artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% emergent vegetation helps in distinguishing their ecology from th ...
s. Many species are limited to very specific habitats and are rare and local. Adults feed in flight and may occur at some distance from their breeding habitat in mixed swarms.


Species

''Somatochlora'' is the most diverse group within the Corduliidae. Of the 42 described species listed below, 25 are North American, 16 are Eurasian, and 1 is
circumboreal The Circumboreal Region in phytogeography is a floristic region within the Holarctic Kingdom in Eurasia and North America, as delineated by such geobotanists as Josias Braun-Blanquet and Armen Takhtajan. It is the largest floristic region in t ...
(''S. sahlbergi''). Two additional fossil species have been assigned to this genus: ''S. brisaci'', based on a wing from the
Upper Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene Epoch made up of two stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma. The ev ...
in
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, and ''S. oregonica'', based on two wings from the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
in
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
.


References

{{Taxonbar, from=Q960357 Corduliidae Anisoptera genera Taxa named by Edmond de Sélys Longchamps