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Snow camouflage is the use of a coloration or pattern for effective
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the ...
in winter, often combined with a different summer camouflage. Summer patterns are typically disruptively patterned combinations of shades of browns and greys, up to black, while winter patterns are dominated by white to match snowy landscapes. Among animals, variable snow camouflage is a type of
seasonal polyphenism A polyphenic trait is a trait for which multiple, discrete phenotypes can arise from a single genotype as a result of differing environmental conditions. It is therefore a special case of phenotypic plasticity. There are several types of polyphen ...
with a distinct winter
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
or
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily #Guard hair, guard hair on top and thick #Down hair, underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as ...
. It is found in birds such as the
rock ptarmigan The rock ptarmigan (''Lagopus muta'') is a medium-sized game bird in the grouse family. It is known simply as the ptarmigan in the UK. It is the official bird for the Canadian territory of Nunavut, where it is known as the ''aqiggiq'' (ᐊᕿ ...
,
lagomorphs The lagomorphs are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae ( hares and rabbits) and the Ochotonidae (pikas). The name of the order is derived from the Ancient Greek ''lagos'' (λα ...
such as the
Arctic hare The Arctic hare (''Lepus arcticus'') is a species of hare highly adapted to living in the Arctic tundra and other icy biomes. The Arctic hare survives with shortened ears and limbs, a small nose, fat that makes up close to 20% of its body, and ...
,
mustelids The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest famil ...
such as the
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
, and one
canid Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...
, the
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small fox native to the Arctic regions of the Northern Hemisphere and common throughout the Arctic tundra biome. It is well adapted to living in co ...
. Since these have evolved separately, the similar appearance is due to
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
. This was used as early evidence for natural selection. Some high Arctic species like the
snowy owl The snowy owl (''Bubo scandiacus''), also known as the polar owl, the white owl and the Arctic owl, is a large, white owl of the true owl family. Snowy owls are native to the Arctic regions of both North America and the Palearctic, breeding mos ...
and
polar bear The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear specie ...
however remain white all year round. In military usage, soldiers often either exchange their disruptively-patterned summer uniforms for thicker snow camouflage uniforms printed with mainly-white versions of
camouflage patterns Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
in winter, or they wear white overalls over their uniforms. Some armies have made use of reversible uniforms, printed in different seasonal patterns on their two sides. Vehicles and guns are often simply repainted in white. Occasionally, aircraft too are repainted in snow camouflage patterns.


Among animals


White as camouflage

Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended fr ...
mentioned the white winter coloration of the ptarmigan in his 1859 ''
Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life''),The book's full original title was ''On the Origin of Species by Me ...
'': The white protective coloration of arctic animals was noted by an early student of camouflage, the naturalist
Alfred Russel Wallace Alfred Russel Wallace (8 January 1823 – 7 November 1913) was a British naturalist, explorer, geographer, anthropologist, biologist and illustrator. He is best known for independently conceiving the theory of evolution through natural se ...
, in his 1889 book ''
Darwinism Darwinism is a scientific theory, theory of Biology, biological evolution developed by the English naturalist Charles Darwin (1809–1882) and others, stating that all species of organisms arise and develop through the natural selection of smal ...
''; he listed the
polar bear The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear specie ...
, the American polar hare, the
snowy owl The snowy owl (''Bubo scandiacus''), also known as the polar owl, the white owl and the Arctic owl, is a large, white owl of the true owl family. Snowy owls are native to the Arctic regions of both North America and the Palearctic, breeding mos ...
and the
gyr falcon The gyrfalcon ( or ) (), the largest of the falcon species, is a bird of prey. The abbreviation gyr is also used. It breeds on Arctic coasts and tundra, and the islands of northern North America and the Eurosiberian region. It is mainly a reside ...
as remaining white all year, while the arctic fox, arctic hare, ermine and ptarmigan change their colour, and observed "the obvious explanation", that it was for concealment, at a time when Darwinism was at a low ebb. Later zoologists such as Hugh B. Cott have echoed his observations, adding that other animals of the Arctic such as
musk ox Musk (Persian: مشک, ''Mushk'') is a class of aromatic substances commonly used as base notes in perfumery. They include glandular secretions from animals such as the musk deer, numerous plants emitting similar fragrances, and artificial sub ...
,
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
,
reindeer Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
,
wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for "gluttony, glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is ...
and
raven A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
never become white "even in the coldest parts of their range". Cott noted that both animals that hunt, like polar bear and stoat, and prey animals like ptarmigan and mountain hare, require camouflage to hide from prey or from predators respectively. There is little experimental evidence for the adaptiveness of white as camouflage, though the ornithologist W. L. N. Tickell, reviewing proposed explanations of white plumage in birds, writes that in the ptarmigan "it is difficult to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite of winter plumage, many Ptarmigan in NE Iceland are killed by Gyrfalcons throughout the winter."


Seasonal polyphenism

Some animals of the far north, like the showshoe and
Arctic hare The Arctic hare (''Lepus arcticus'') is a species of hare highly adapted to living in the Arctic tundra and other icy biomes. The Arctic hare survives with shortened ears and limbs, a small nose, fat that makes up close to 20% of its body, and ...
s,
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small fox native to the Arctic regions of the Northern Hemisphere and common throughout the Arctic tundra biome. It is well adapted to living in co ...
,
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
, and
rock ptarmigan The rock ptarmigan (''Lagopus muta'') is a medium-sized game bird in the grouse family. It is known simply as the ptarmigan in the UK. It is the official bird for the Canadian territory of Nunavut, where it is known as the ''aqiggiq'' (ᐊᕿ ...
change their coat colour (by moulting and growing new fur or feathers) from brown or grey summer
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the ...
to white in the winter; the Arctic fox is the only species in the dog family to do so. However, Arctic hares which live in the far north of
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, where summer is very short, remain white year-round. Since these animals in widely separated groups have evolved separately, the similarity of coloration is due to
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
, on the presumption that
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Charle ...
favours a particular coloration in a particular environment. The
seasonal polyphenism A polyphenic trait is a trait for which multiple, discrete phenotypes can arise from a single genotype as a result of differing environmental conditions. It is therefore a special case of phenotypic plasticity. There are several types of polyphen ...
in
willow grouse The willow ptarmigan () (''Lagopus lagopus'') is a bird in the grouse subfamily Tetraoninae of the pheasant family Phasianidae. It is also known as the willow grouse and in Ireland and Britain, where the subspecies '' L. l. scotica'' was previ ...
differs between Scottish and Scandinavian populations. In Scotland, grouse have two plumages (breeding and non-breeding), while in Scandinavia there is a third plumage, a white winter morph. The genetic basis for this is not in the
melanin Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino ...
pigment system, and is probably due to regulatory changes. The behaviour of moulting females in springtime depends on their plumage state: they tend to sit on snow while they are mainly white, but choose the border between bare ground and snow when they have more dark feathers. They seem to be choosing the best compromise between camouflage and food quality.
Climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
from
global warming In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
is leading to a mismatch between the seasonal coat coloration of arctic animals such as
snowshoe hare The snowshoe hare (''Lepus americanus''), also called the varying hare or snowshoe rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet. The animal's feet prevent it from sink ...
s with the increasingly snow-free landscape.


Military usage

The principle of varying coloration with the changing seasons has military applications.


First World War

In the
First World War World War I (28 July 1914 11 November 1918), often abbreviated as WWI, was one of the deadliest global conflicts in history. Belligerents included much of Europe, the Russian Empire, the United States, and the Ottoman Empire, with fightin ...
, firing and observation positions were hand-painted in disruptive patterns by artists known as
camoufleurs A camoufleur or camouflage officer is a person who designed and implemented military camouflage in one of the world wars of the twentieth century. The term originally meant a person serving in a First World War French military camouflage unit. In ...
, and they sometimes varied their patterns seasonally. Uniforms were largely of a single colour, such as the British
khaki The color khaki (, ) is a light shade of tan with a slight yellowish tinge. Khaki has been used by many armies around the world for uniforms and equipment, particularly in arid or desert regions, where it provides camouflage relative to sandy ...
; but snow camouflage clothes came into use in some armies by 1917. For example, Austro-Hungarian troops on the Italian front used
ski A ski is a narrow strip of semi-rigid material worn underfoot to glide over snow. Substantially longer than wide and characteristically employed in pairs, skis are attached to ski boots with ski bindings, with either a free, lockable, or partial ...
s and wore snow camouflage smocks and overtrousers over their
uniforms A uniform is a variety of clothing worn by members of an organization while participating in that organization's activity. Modern uniforms are most often worn by armed forces and paramilitary organizations such as police, emergency services, ...
, and improvised white cloths over their uniform caps.


Second World War

Several armies in the
Second World War World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposin ...
in Northern European countries preferred separate winter uniforms rather than oversuits. The
Waffen-SS The (, "Armed SS") was the combat branch of the Nazi Party's ''Schutzstaffel'' (SS) organisation. Its formations included men from Nazi Germany, along with Waffen-SS foreign volunteers and conscripts, volunteers and conscripts from both occup ...
went a step further, developing reversible uniforms with separate schemes for summer and autumn, as well as white winter oversuits. Other German units fighting in Eastern Europe were at first poorly equipped for winter, having to make do with ordinary summer uniforms, but in the winter of 1942 to 1943 new white two-piece hooded oversuits with long mitten gauntlets started to arrive. American troops in Europe in the winter of 1944 to 1945 improvised snow capes and helmet covers from white cloth such as bed linen. The Red Army issued a report, "Tactical and Technical trends, No. 17" in January 1943 on the camouflage of tanks in winter. It advised either all-white using zinc white or titanium white paint for level, open country, or disruptive two-colour winter camouflage for areas with more variety including "forests, underbrush, small settlements, thawed patches of earth". The two colours could be achieved either by leaving around a quarter to a third of the vehicle's summer camouflage uncovered, or by repainting the whole vehicle in white with dark gray or gray-brown spots. Units were advised not to paint all their vehicles identically, but to have some tanks all white, some in white with green stripes, and some in white and gray or gray-brown. Winter camouflage was not limited to paint: tracks left in the snow were to be obliterated, vehicles parked in cover, headlights covered with white fabric, shelters constructed, or else vehicles covered with white fabric, or dark fabric scattered with snow.


Post-war

Later in the 20th century, as disruptively patterned uniforms became widespread, winter variants of patterns such as ''
Flecktarn ''Flecktarn'' (; "mottled camouflage"; also known as ''Flecktarnmuster'' or ''Fleckentarn'') is a family of 3-, 4-, 5- or 6-color disruptive camouflage patterns, the most common being the five-color pattern, consisting of dark green, light green ...
'' were developed, the background colour (such as green) being replaced with white to form a ''Schneetarn'' (snow camouflage pattern). Aircraft deployed in winter have sometimes been snow camouflaged, as with
Sepecat Jaguar The SEPECAT Jaguar is an Anglo-French jet attack aircraft originally used by the British Royal Air Force and the French Air Force in the close air support and nuclear strike role. It is still in service with the Indian Air Force. Originall ...
s on exercise in Norway. Snow camouflage is manufactured commercially for use by
hunters Hunting is the human practice of seeking, pursuing, capturing, or killing wildlife or feral animals. The most common reasons for humans to hunt are to harvest food (i.e. meat) and useful animal products (fur/ hide, bone/tusks, horn/antler, et ...
. For example, NaturalGear derive their patterns from photographic images using "the colors of nature".


References

{{Camouflage Camouflage Animal coat colors Military camouflage