Snouted Cobra
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The snouted cobra (''Naja annulifera''), also called the banded Egyptian cobra, is a highly venomous species of cobra found in Southern Africa.


Description

The snouted cobra is a relatively large species. Adult specimens average between in length, but they may reach lengths of . Colouration of dorsal scales may vary from yellowish to greyish-brown, dark brown or blue-black. Ventral scale colouration is yellow with darker mottles. A banded phase occurs throughout the species' range and is blue-black with 7-11 yellow to yellow-brown cross bars, the lighter bands being half the width of the darker bands. The latter colour phase is more common in males. Ventrally, it is yellow mottled with black. A darker throat band is present and is usually more prominent in juveniles.


Scalation

Midbody scales are in 19 rows (rarely 21) with 175-203 ventrals. There are 51-65 paired subcaudals and the anal shield is entire. There are seven (sometimes eight) upper labials that do not enter the eye, eight or 9 (rarely 10) lower labials, as well as one preocular (sometimes two) and two (sometimes one or three) postoculars. Temporals are variable.


Distribution

This species is found in northeast
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, southern
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...
, eastern
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label=Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalahar ...
,
Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northeast ...
, throughout
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and Mozam ...
, and parts of
Eswatini Eswatini ( ; ss, eSwatini ), officially the Kingdom of Eswatini and formerly named Swaziland ( ; officially renamed in 2018), is a landlocked country in Southern Africa. It is bordered by Mozambique to its northeast and South Africa to its no ...
.


Habitat and ecology

Snouted cobras inhabit arid and moist
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
, particularly in
bushveld The Bushveld (from af, bosveld, af, bos 'bush' and af, veld) is a tropical and subtropical grasslands, savannas, and shrublands, sub-tropical woodland ecoregion of Southern Africa. It encompasses most of Limpopo Province and a small part of ...
and lowveld areas. It is not found in forests. As a large cobra, it often has a permanent home base or lair in an abandoned termite mound, where it will reside for years if left undisturbed. It is a nocturnal species, foraging for food from dusk onwards. It enjoys basking in the sun during the day near its lair or retreat. This species can be quite nervous and will strike to defend itself if threatened. Like other cobras, when disturbed, it usually raises the front-third of its body when extending its hood and hissing. Very large adults are able to lift as much as 0.5 m of the body off the ground while spreading a wide, impressive hood. However, given the opportunity, it will escape to the nearest hole or crevice. Like the
rinkhals The rinkhals (; ''Hemachatus haemachatus''), also known as the ringhals or ring-necked spitting cobra, is a species of venomous snake in the family Elapidae. The species is found in parts of southern Africa. It is not a true cobra in that it do ...
, it may sham death if threatened, but this is rare. It preys on
toads Toad is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands. A distinction between frogs and toads is not made in scientif ...
,
rodents Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are nat ...
,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s and their eggs,
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s and other snakes, especially puff adders (''
Bitis arietans The puff adder (''Bitis arietans'') is a viper species found in savannahs and grasslands from Morocco and western Arabia throughout Africa except for the Sahara and rainforest regions.U.S. Navy. 1991. ''Venomous Snakes of the World''. US Govt. ...
''). It often raids poultry runs and can become a nuisance. It is preyed upon by
birds of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators ...
and other snakes.


Reproduction

This is an
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
species, laying between 8 and 33 eggs in early summer. The young average in length.


Taxonomy

It was formerly considered a subspecies of the
Egyptian cobra The Egyptian cobra (''Naja haje''), also known as Ouraeus ''(derived from the Ancient Greek word: ''οὐραῖος'' - )'', is one of the most venomous snakes in North Africa, which has caused many snakebite incidents to humans. It averages rou ...
(''Naja haje''), as was
Anchieta's cobra Anchieta's cobra (''Naja anchietae''), sometimes referred to as the Angolan cobra, is a species of venomous snake in the family Elapidae. The species is native to Southern Africa. Taxonomy Anchieta's cobra was first described by Portuguese zool ...
(''Naja anchietae''). The latter taxon was subsequently considered to be a subspecies of the snouted cobra, before being split off as a distinct species.


Venom

It is a highly venomous species with
neurotoxic Neurotoxicity is a form of toxicity in which a biological, chemical, or physical agent produces an adverse effect on the structure or function of the central and/or peripheral nervous system. It occurs when exposure to a substance – specificall ...
venom. Intravenous value is 1.98 mg/kg. A bite can affect breathing, and if left untreated, may cause respiratory failure and death. Initial symptoms include pain and local swelling that may result in blistering. Typically, victims are bitten on the lower leg, usually at night.


References

{{DEFAULTSORT:cobra, snouted
snouted cobra The snouted cobra (''Naja annulifera''), also called the banded Egyptian cobra, is a highly venomous species of cobra found in Southern Africa. Description The snouted cobra is a relatively large species. Adult specimens average between in len ...
Snakes of Africa Reptiles of Southern Africa
snouted cobra The snouted cobra (''Naja annulifera''), also called the banded Egyptian cobra, is a highly venomous species of cobra found in Southern Africa. Description The snouted cobra is a relatively large species. Adult specimens average between in len ...
snouted cobra The snouted cobra (''Naja annulifera''), also called the banded Egyptian cobra, is a highly venomous species of cobra found in Southern Africa. Description The snouted cobra is a relatively large species. Adult specimens average between in len ...