The small angle shades (''Euplexia lucipara'') is a

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

of the family

Noctuidae The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family of moths. They are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly changing, along with the other f ...

. It is distributed throughout the

Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...

. The species was first described by

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

in his 1758 10th edition of ''Systema Naturae''. As the common name suggests, this species is closely related to the

angle shades The angle shades (''Phlogophora meticulosa'') is a moth of the family Noctuidae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. It is distributed throughout Europe as far east as the Urals and a ...

(''Phlogophora meticulosa''), and is considerably smaller (

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...

30–35 mm), but does not especially resemble that species. The forewings are dark brown with a broad, pale subterminal band, wider and paler towards the

costa Costa may refer to: Biology * Rib (Latin: ''costa''), in vertebrate anatomy * Costa (botany), the central strand of a plant leaf or thallus * Costa (coral), a stony rib, part of the skeleton of a coral * Costa (entomology), the leading edge of th ...

. The hindwings are whitish at the base, graduating to brown at the margins. In the

British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...

this species flies at night in June and July, with a second generation sometimes emerging in September. It is attracted to light and sugar.

Distribution

It is found throughout Europe, in

Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...

, in

western Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...

and through the

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

, and

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

Technical description

Forewing rufous ochreous tinged with purplish and mixed with olive brown; median area, a triangular blotch on inner margin near base, and a narrow praesubmarginal cloud deep olive; costa to beyond middle, the terminal area, and the orbicular stigma leaden purple; lines indistinct, the outer and inner approximating on inner margin; claviform stigma olive, dark edged; orbicular roundish, oblique, with paler ring; reniform conspicuously whitish, containing a double brown lunule; submarginal line pale, waved; veins towards termen dotted dark and pale; hindwing ochreous with dark cell mark and veins, darker in terminal half where there is visible a dark pale-edged outer and submarginal line.

Biology

Larva velvety green with darker green oblique subdorsal stripes; tubercles pale; spiracular line yellowish white; 11th segment slightly swollen, with a pair of white dots; head green. The

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...

e often feed on

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

s and the species is usually associated with them. Other recorded food plants include

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...

, false bindweed,

dogwood ''Cornus'' is a genus of about 30–60 species of woody plants in the family Cornaceae, commonly known as dogwoods, which can generally be distinguished by their blossoms, berries, and distinctive bark. Most are deciduous trees or shrub ...

, larkspur,

willowherb Willowherbs are annual or perennial herbaceous plants in the flowering plant family Onagraceae. Willowherb may refer to: * A species of willowherb in the genus ''Epilobium'' * A species of willowherb in the genus ''Chamaenerion ''Chamaenerion'' ...

ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non- gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...

ivy ''Hedera'', commonly called ivy (plural ivies), is a genus of 12–15 species of evergreen climbing or ground-creeping woody plants in the family Araliaceae, native to western, central and southern Europe, Macaronesia, northwestern Africa and ...

lettuce Lettuce (''Lactuca sativa'') is an annual plant of the family Asteraceae. It is most often grown as a leaf vegetable, but sometimes for its stem and seeds. Lettuce is most often used for salads, although it is also seen in other kinds of food, ...

privet A privet is a flowering plant in the genus ''Ligustrum''. The genus contains about 50 species of erect, deciduous or evergreen shrubs, sometimes forming small or medium-sized trees, native species, native to Europe, north Africa, Asia, many in ...

, loosestrife,

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

buttercup ''Ranunculus'' is a large genus of about almost 1700 to more than 1800 species of flowering plants in the family Ranunculaceae. Members of the genus are known as buttercups, spearworts and water crowfoots. The genus is distributed in Europe, ...

, currant,

raspberry The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with w ...

willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...

tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...

, coltsfoot,

nettle {{redirect, Nettle Nettle refers to plants with stinging hairs, particularly those of the genus ''Urtica''. It can also refer to plants which resemble ''Urtica'' species in appearance but do not have stinging hairs. Plants called "nettle" include ...

, and

guelder rose ''Viburnum opulus'', the guelder-rose or guelder rose () is a species of flowering plant in the family Adoxaceae (formerly Caprifoliaceae) native to Europe, northern Africa and central Asia. Description ''Viburnum opulus'' is a deciduous shru ...

. The species overwinters as

pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...

Gallery

Image:Britishentomologyvolume5Plate619.jpg, Illustration from ''

British Entomology ''British Entomology'' is a classic work of entomology by John Curtis, FLS. It is subtitled ''Being Illustrations and Descriptions of the Genera of Insects found in Great Britain and Ireland: Containing Coloured Figures from Nature of the Most ...

'', Volume 5, by John Curtis Image:Euplexia lucipara, Small Angleshades, Flint, North Wales, Aug 2017 (36443636021).jpg, Larva.Flint, North Wales

Notes

References

Chinery, Michael Michael Chinery (born 1938, in London) is an English naturalist. He studied in Cambridge where he graduated in natural sciences and anthropology. He edits '' Cecidology'', the journal of the British Plant Gall Society The British Plant Gall ...

(1986, reprinted 1991). ''Collins Guide to the Insects of Britain and Western Europe''. * Skinner, Bernard ''

The Colour Identification Guide to Moths of the British Isles ''The Colour Identification Guide to Moths of the British Isles (Macrolepidoptera)'' by Bernard Skinner is a single volume identification guide to the macromoths of Britain and Ireland published by Viking Books, often referred by moth recorders ...

''. 1984

External links

* * Taxonomy
''Lepiforum e.V.''''De Vlinderstichting''
{{Taxonbar , from=Q649260 Euplexia Moths described in 1758 Moths of Africa Moths of Asia Moths of Europe Moths of Japan Taxa named by Carl Linnaeus