Skywalker Hoolock Gibbon
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The Skywalker hoolock gibbon or Gaoligong hoolock gibbon (''Hoolock tianxing'') is an arboreal
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
in the
gibbon Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical rainforest from eastern Bangladesh to Northeast India ...
family,
Hylobatidae Gibbons () are apes in the family Hylobatidae (). The family historically contained one genus, but now is split into four extant genera and 20 species. Gibbons live in subtropical and tropical rainforest from eastern Bangladesh to Northeast India ...
. It is one of three species of
hoolock gibbon The hoolock gibbons are three primate species of genus ''Hoolock'' in the gibbon family, Hylobatidae, native to eastern Bangladesh, Northeast India, Myanmar, and Southwest China. Description Hoolocks are the second-largest of the gibbons, after ...
and was first described in January 2017 in the ''
American Journal of Primatology The ''American Journal of Primatology'' is a monthly peer-reviewed scientific journal and the official journal of the American Society of Primatologists. It was established in 1981 and covers all areas of primatology, including the behavioral ecol ...
''. The Skywalker hoolock gibbon is one of two species of Eastern hoolocks: ''H. tianxing'' and ''H. leuconedys''. Researchers estimate ''H. tianxing'' diverged from ''H. leuconedys'' roughly 490,000 years ago. The Eastern hoolock is vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
, with a population of 310,000-370,000 individuals. Of this population, ''H. tianxing'' makes up less than 150 individuals, making the Skywalker hoolock gibbon an endangered species.


Etymology

The Skywalker hoolock gibbon is named after
Luke Skywalker Luke Skywalker is a fictional character and the protagonist of the original film trilogy of the ''Star Wars'' franchise created by George Lucas. Portrayed by Mark Hamill, Luke first appeared in ''Star Wars'' (1977), and he returned in ''The ...
from the
Star Wars ''Star Wars'' is an American epic film, epic space opera multimedia franchise created by George Lucas, which began with the Star Wars (film), eponymous 1977 film and quickly became a worldwide popular culture, pop-culture Cultural impact of S ...
franchise, as the scientists who described it are fans of the franchise. The specific name ''tianxing'' is the
Pinyin Hanyu Pinyin (), often shortened to just pinyin, is the official romanization system for Standard Mandarin Chinese in China, and to some extent, in Singapore and Malaysia. It is often used to teach Mandarin, normally written in Chinese for ...
transliteration Transliteration is a type of conversion of a text from one writing system, script to another that involves swapping Letter (alphabet), letters (thus ''wikt:trans-#Prefix, trans-'' + ''wikt:littera#Latin, liter-'') in predictable ways, such as ...
of the
Chinese Chinese can refer to: * Something related to China * Chinese people, people of Chinese nationality, citizenship, and/or ethnicity **''Zhonghua minzu'', the supra-ethnic concept of the Chinese nation ** List of ethnic groups in China, people of va ...
terms 天 (''tiān'') meaning "heaven" and 行 (''xíng'') meaning "movement". The name is a reference to
brachiation Brachiation (from "brachium", Latin for "arm"), or arm swinging, is a form of arboreal locomotion in which primates swing from tree limb to tree limb using only their arms. During brachiation, the body is alternately supported under each forelimb ...
, the main locomotory mode of gibbons, and derived from the text of the ''
I Ching The ''I Ching'' or ''Yi Jing'' (, ), usually translated ''Book of Changes'' or ''Classic of Changes'', is an ancient Chinese divination text that is among the oldest of the Chinese classics. Originally a divination manual in the Western Zho ...
'', an ancient Chinese book of divination.


External Morphology

The external morphological traits of the Skywalker hoolock gibbon differ from those of the ''H. leuconedys''. The ''H. tianxing'' has distinguished white eyebrows, and their eyebrow streaks are thinner and separated by a larger gap than those of the ''H. leuconedys''. They have large beards that are either all black or brown, rather than the white beards of the ''H. leuconedys'', and they do not have pieces of white hair on their suborbital area. The ''H. tianxing'' have tufts of fur on their genital areas that are coloured either black, brown or dark grey, compared to the white tufts on the genital areas of ''H. leuconedys''.
Sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
is present in Skywalker hoolock gibbons. In the adult male ''H. tianxing'', their ventral
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily #Guard hair, guard hair on top and thick #Down hair, underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as ...
is coloured dark brown, and their dorsal
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily #Guard hair, guard hair on top and thick #Down hair, underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as ...
has a brownish overlay. In the adult female ''H. tianxing'', their ventral
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily #Guard hair, guard hair on top and thick #Down hair, underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as ...
is coloured yellow/white or reddish blonde. Females also have distinctive incomplete white face rings, with sparse hairs on their lateral orbital and suborbital regions. As they get older, the colouring of their genital tufts grows fainter and lighter. Juveniles of the species can be identified by their lack of white fur on their chin or under their eyes.


Habitat/Distribution

The Skywalker hoolock gibbon can be found in the
montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
forests of eastern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
and southwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
in the Mt. Gaoligong region, located between the
Salween River , ''Mae Nam Salawin'' ( , name_etymology = , image = Sweet_View_of_Salween_River_in_Tang_Yan_Township,_Shan_State,_Myanmar.jpg , image_size = , image_caption = Salween River in Shan State, Myanmar , map ...
and the Nmai tributary of the
Irrawaddy River The Irrawaddy River ( Ayeyarwady River; , , from Indic ''revatī'', meaning "abounding in riches") is a river that flows from north to south through Myanmar (Burma). It is the country's largest river and most important commercial waterway. Origi ...
. Hoolocks were first recorded in this region in 1917,  and this is the easternmost habitat of any hoolock species. Most of the ''H. tianxing'' population can be found in unprotected forests, and they live in small patches of forest with very rough terrain. 11 solitary and 32 groups of Skywalker hoolock gibbons have been recorded in the wild, but their total population is composed of less than 150 individuals.


Behaviour

Skywalker hoolock gibbons sleep in tall trees, on the thinner branches near the crown of the tree. They choose their sleeping tree for five different reasons:
foraging Foraging is searching for wild food resources. It affects an animal's Fitness (biology), fitness because it plays an important role in an animal's ability to survive and reproduce. Optimal foraging theory, Foraging theory is a branch of behaviora ...
abilities, range or resource defence abilities,
thermoregulation Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
abilities, comfort and hygiene, and antipredation reasons. They enter their sleeping trees before sunset in order to avoid
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
predators who are very active at that time, and once their tree has been selected, they move to it fast and directly before settling quietly for the night. They rarely spend consecutive nights sleeping in the same tree. During the cold season in
Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...
province, from November to March, the ''H. tianxing'' choose sleeping trees closer to food trees and at a lower elevation where it is warmer, and choose to sleep longer and huddle together to
thermoregulate Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperature ...
. Gibbons are known for having loud songs, specific to their species. They usually perform these songs from their sleeping trees just after dawn, and these songs last an average of 22.5 minutes.


Breeding

There are four steps to the
mating In biology, mating is the pairing of either opposite-sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reproduc ...
ritual of a Skywalker hoolock gibbon.Huang, X., Zhou, W., & Ai, H. (2010). Mating behavior of Hoolock gibbon (Hoolock hoolock) in the field: a case study at Mt. Gaoligong, Yunnan, China. ''Journal of Southwest Forestry University'', ''30''(2), 52-55. First, the females raise their
buttocks The buttocks (singular: buttock) are two rounded portions of the exterior anatomy of most mammals, located on the posterior of the pelvic region. In humans, the buttocks are located between the lower back and the perineum. They are composed ...
as an invitation for the male to begin mating. To accept this request, the male ''H. tianxing'' approaches the female. The male mounts the female for mating, and they have a period of rest after the mating has concluded. In the wild, it is the female ''H. tianxing'' that dominates this ritual.


Diet

The Skywalker hoolock gibbon is known to consume 36 different botanic species. Their preference is always the
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
of these trees, followed by the
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
, and opting for the stems and
flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
last. Female ''H. tianxing'' are reported to eat more than the males, especially during
spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a ...
and
autumn Autumn, also known as fall in American English and Canadian English, is one of the four temperate seasons on Earth. Outside the tropics, autumn marks the transition from summer to winter, in September ( Northern Hemisphere) or March ( Sou ...
.


Threats

The Skywalker hoolock gibbon is an
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
species, with less than 150 individuals reported in the wild. This population is fragmented between different forest areas, causing problems for further population decline due to the small population size and high likelihood of isolation. They are threatened by an abundance of
illegal hunting Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...
and
trading Trade involves the transfer of goods and services from one person or entity to another, often in exchange for money. Economists refer to a system or network that allows trade as a market. An early form of trade, barter, saw the direct exchan ...
of the species, and their
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
is prone to destruction for the use of
cardamon Cardamom (), sometimes cardamon or cardamum, is a spice made from the seeds of several plants in the genera ''Elettaria'' and ''Amomum'' in the family Zingiberaceae. Both genera are native to the Indian subcontinent and Indonesia. They are rec ...
cultivation. In China, 22 hoolocks are in
captivity Captivity, or being held captive, is a state wherein humans or other animals are confined to a particular space and prevented from leaving or moving freely. An example in humans is imprisonment. Prisoners of war are usually held in captivity by a ...
, but only two of these are ''H. tianxing''. No females are currently in captivity, making
reproduction Reproduction (or procreation or breeding) is the biological process by which new individual organisms – "offspring" – are produced from their "parent" or parents. Reproduction is a fundamental feature of all known life; each individual or ...
in captivity impossible.


References

{{Taxonbar, from=Q28197046
Skywalker hoolock gibbon The Skywalker hoolock gibbon or Gaoligong hoolock gibbon (''Hoolock tianxing'') is an arboreal primate in the gibbon family, Hylobatidae. It is one of three species of hoolock gibbon and was first described in January 2017 in the ''American Jour ...
Primates of East Asia Mammals of Myanmar Fauna of Yunnan
Skywalker hoolock gibbon The Skywalker hoolock gibbon or Gaoligong hoolock gibbon (''Hoolock tianxing'') is an arboreal primate in the gibbon family, Hylobatidae. It is one of three species of hoolock gibbon and was first described in January 2017 in the ''American Jour ...
Star Wars Endangered fauna of Asia