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The skuas are a group of predatory
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s with seven species forming the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''Stercorarius'', the only genus in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Stercorariidae. The three smaller skuas, the
long-tailed skua The long-tailed skua or long-tailed jaeger (''Stercorarius longicaudus'') is a seabird in the skua family Stercorariidae. Etymology The word "jaeger" is derived from the German word ''Jäger'', meaning "hunter". The English word "skua" comes ...
, the
Arctic skua The parasitic jaeger (''Stercorarius parasiticus''), also known as the Arctic skua, Arctic jaeger or parasitic skua, is a seabird in the skua family Stercorariidae. It is a migratory species that breeds in Northern Scandinavia, Scotland, Ice ...
, and the pomarine skua are called jaegers in
North American English North American English (NAmE, NAE) is the most generalized variety (linguistics), variety of the English language as spoken in the United States and Canada. Because of their related histories and cultures, plus the similarities between the pron ...
. The English word "skua" comes from the Faroese name for the
great skua The great skua (''Stercorarius skua''), sometimes known by the name bonxie in Britain, is a large seabird in the skua family Stercorariidae. It is roughly the size of a herring gull. It mainly eats fish caught at the sea surface or taken f ...
, , with the island of
Skúvoy Skúvoy or Skúgvoy ( da, Skuø) is an island in the central Faroe Islands, located to the south of Sandoy. It is named after the large number of great skua present on the island (who have a habit of attacking intruders). There is only one s ...
renowned for its colony of that bird. The general Faroese term for skuas is . The word "jaeger" is derived from the
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ger ...
word , meaning "hunter". The genus name ''Stercorarius'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
and means "of dung"; because the food disgorged by other birds when pursued by skuas was once thought to be excrement. Skuas nest on the ground in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
and Arctic regions, and are long-distance
migrants Migrant may refer to: Human migration *Human migration *Emigration, leaving one's resident country with the intent to settle elsewhere *Immigration, movement into a country with the intent to settle * Economic migrant, someone who emigrates from o ...
. They have even been sighted at the
South Pole The South Pole, also known as the Geographic South Pole, Terrestrial South Pole or 90th Parallel South, is one of the two points where Earth's axis of rotation intersects its surface. It is the southernmost point on Earth and lies antipod ...
.


Biology and habits

Outside the breeding season, skuas take fish,
offal Offal (), also called variety meats, pluck or organ meats, is the organs of a butchered animal. The word does not refer to a particular list of edible organs, which varies by culture and region, but usually excludes muscle. Offal may also ref ...
, and
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
. Many practice
kleptoparasitism Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when fo ...
, which comprises up to 95% of the feeding methods of wintering skuas, by chasing gulls, terns and other seabirds to steal their catches, regardless of the size of the species attacked (up to three times heavier than the attacking skua). The larger species, such as the
great skua The great skua (''Stercorarius skua''), sometimes known by the name bonxie in Britain, is a large seabird in the skua family Stercorariidae. It is roughly the size of a herring gull. It mainly eats fish caught at the sea surface or taken f ...
, also regularly kill and eat adult birds, such as
puffin Puffins are any of three species of small alcids ( auks) in the bird genus ''Fratercula''. These are pelagic seabirds that feed primarily by diving in the water. They breed in large colonies on coastal cliffs or offshore islands, nesting in c ...
s and
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, ...
s, and have been seen killing birds as large as a
grey heron The grey heron (''Ardea cinerea'') is a long-legged wading bird of the heron family, Ardeidae, native throughout temperate Europe and Asia and also parts of Africa. It is resident in much of its range, but some populations from the more northern ...
. On the breeding grounds, the three, more slender northern breeding species commonly eat
lemming A lemming is a small rodent, usually found in or near the Arctic in tundra biomes. Lemmings form the subfamily Arvicolinae (also known as Microtinae) together with voles and muskrats, which form part of the superfamily Muroidea, which also incl ...
s. Those species that breed in the southern oceans largely feed on fish that can be caught near their colonies. The eggs and chicks of other seabirds, primarily
penguin Penguins ( order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adapt ...
s, are an important food source for most skua species during the nesting season. In the southern oceans and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest co ...
region, some skua species (especially the
south polar skua The south polar skua (''Stercorarius maccormicki'') is a large seabird in the skua family, Stercorariidae. An older name for the bird is MacCormick's skua, after explorer and naval surgeon Robert McCormick, who first collected the type speci ...
) will readily scavenge carcasses at breeding colonies of both
penguin Penguins ( order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adapt ...
s and
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammal, marine mammals. They comprise the extant taxon, extant family (biology ...
s. Skuas will also kill live penguin chicks. In these areas, the skuas will often forfeit their catches to the considerably larger and very aggressive giant petrels. Skuas have also been observed to directly pilfer
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digest solid food. Immune factors and immune-modulati ...
from the elephant seal's teat. They are medium to large
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, typically with grey or brown plumage, often with white markings on the wings. The skuas range in size from the
long-tailed skua The long-tailed skua or long-tailed jaeger (''Stercorarius longicaudus'') is a seabird in the skua family Stercorariidae. Etymology The word "jaeger" is derived from the German word ''Jäger'', meaning "hunter". The English word "skua" comes ...
, ''Stercorarius longicauda'', at , to the brown skua, ''Stercorarius antarcticus'', at . On average, a skua is about long, and across the wings. They have longish bills with a hooked tip, and webbed feet with sharp claws. They look like large dark gulls, but have a fleshy cere above the upper mandible. The skuas are strong, acrobatic fliers. They are generally aggressive in disposition. Potential predators approaching their nests will be quickly attacked by the parent birds, which usually target the heads of intruders – a practice known as ' divebombing'.


Taxonomy

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''Stercorarius'' was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published wo ...
in 1760 with the
parasitic jaeger The parasitic jaeger (''Stercorarius parasiticus''), also known as the Arctic skua, Arctic jaeger or parasitic skua, is a seabird in the skua family Stercorariidae. It is a migratory species that breeds in Northern Scandinavia, Scotland, Iceland, ...
(''Stercorarius parasiticus'') as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
. Skuas are related to
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, ...
s,
wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
s, auks, and
skimmer Skimmer may refer to: Animals *Skimmer (bird), a common name for birds in the genus ''Rynchops'' *Skimmer (dragonfly), a common name for dragonflies in the family Libellulidae *Water strider or skimmer, a common name for insects in the family Ge ...
s. In the three smaller species, all nesting exclusively in the Holarctic, breeding adults have the two central tail feathers obviously elongated, and at least some adults have white on the underparts and pale yellow on the neck. These characteristics are not shared by the larger species, all native to the Southern Hemisphere except for the great skua. Therefore, the skuas are often split into two genera, with only the smaller species retained in ''Stercorarius'', and the large species placed in ''Catharacta''. However, based on
genetics Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinian friar worki ...
, behavior, and feather lice, the overall relationship among the species is best expressed by placing all in a single genus. The pomarine and great skuas' mitochondrial DNA (inherited from the mother) is in fact more closely related to each other than it is to either Arctic or long-tailed skuas, or to the Southern Hemisphere species.Cohen, Baker, Belchschmidt, Dittmann, Furness, Gerwin, Helbig, de Korte, Marshall, Palma, Peter, Ramli, Siebold, Willcox, Wilson and Zink (1997)
Enigmatic phylogeny of skuas.
''Proc. Biol. Sci.'' 264(1379):181–190.
Thus, hybridization must have played a considerable role in the evolution of the diversity of Northern Hemisphere skuas.


Species

The genus contains seven species:


References


External links

*
Skua videos
on the Internet Bird Collection {{Authority control Seabirds Taxa named by Mathurin Jacques Brisson