Simoedosaurus Lemoinei Cernay
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''Simoedosaurus'' is an extinct reptile known from the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
of North America, Europe and western Asia, and a member of the
Choristodera Choristodera (from the Greek χωριστός ''chōristos'' + δέρη ''dérē'', 'separated neck') is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the late Miocene (168 to 1 ...
, a group of aquatic reptiles that lived in the Northern Hemisphere from the Jurassic to the early Cenozoic. Described other species, ''S. dakotensis'' later got its own genus, ''
Kosmodraco ''Kosmodraco'' is a genus of large bodied choristodere from the Paleocene of North America. Originally described as a species of ''Simoedosaurus'', it was found to represent a distinct genus in 2022. Multiple fossil skulls show a relatively short ...
''.


Taxonomy

French paleontologist
Paul Gervais Paul Gervais full name François Louis Paul Gervais (26 September 1816 – 10 February 1879) was a French palaeontologist and entomologist. Biography Gervais was born in Paris, where he obtained the diplomas of doctor of science and of medicine ...
described ''Simoedosaurus'' in 1877. Though similar to and contemporaneous, ''Simoedosaurus'' is not closely related to the North American ''Champsosaurus'', instead it appears to be most closely related to '' Tchoiria'' and ''
Ikechosaurus ''Ikechosaurus'' is an extinct genus of choristodere reptile which existed in China and Mongolia during the Early Cretaceous. It contains the species ''Ikechosaurus sunailinae'' and ''Ikechosaurus gaoi''. It belongs to the crocodilian-like clade ...
'' from the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
of Asia. It therefore may represent a species that immigrated into North America from Asia in the wake of the Cretaceous-Tertiary mass extinction event, though the absence of choristoderes in the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
of Asia makes this merely a paleogeographical speculation.


Biology

''Simoedosaurus'' was an aquatic predator, specialised to a fully aquatic lifestyle; though ''
Champsosaurus ''Champsosaurus'' is an extinct genus of crocodile-like choristodere reptile, known from the Late Cretaceous and early Paleogene periods of North America and Europe (Campanian-Paleocene). The name ''Champsosaurus'' is thought to come from , () sa ...
'' might have still come ashore to lay eggs,
ovovivipary Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
is known in other choristoderes. It in particular possesses broader, stronger jaws than other longirostrine choristoderes, including its closest relatives, suggesting that it was capable of tackling larger prey. ''Simoedosaurus'' does occur in sites where aquatic crocodilians are present, including brevirostrine forms like ''
Borealosuchus ''Borealosuchus'' (meaning "boreal crocodile") is an extinct genus of crocodyliforms that lived from the Late Cretaceous to the Eocene in North America. It was named by Chris Brochu in 1997 for several species that had been assigned to ''Leidyos ...
''; the extent of competition between both groups, if there was any, is still unresolved. Like other neo-choristoderes it has nasal conchae, suggesting it could regulate its own body temperature, explaining its ability to live in cold waters.


Range

The earliest records of ''Simoedosaurus'' are from the Early Paleocene (Puercan Land Mammal Age) of Saskatchewan. It persisted until the Late Palaecene in North America, and has also been found in the Late Paleocene of France. The youngest remains seem to occur in the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
.


References

{{Taxonbar, from=Q2164813 Choristodera Paleocene reptiles Eocene reptiles Extinct animals of Europe Extinct animals of North America Prehistoric marine reptiles Prehistoric reptile genera