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The short-eared owl (''Asio flammeus'') is a widespread grassland species in the family Strigidae. Owls belonging to genus ''
Asio ''Asio'' is a genus of typical owls, or true owls, in the family Strigidae. This group has representatives over most of the planet, and the short-eared owl is one of the most widespread of all bird species, breeding in Europe, Asia, North an ...
'' are known as the eared owls, as they have tufts of feathers resembling mammalian ears. These "ear" tufts may or may not be visible. The short-eared owl will display its tufts when in a defensive pose, although its very short tufts are usually not visible. The short-eared owl is found in open country and grasslands.


Taxonomy

The short-eared owl was formally described in 1763 by the
Lutheran Lutheranism is one of the largest branches of Protestantism, identifying primarily with the theology of Martin Luther, the 16th-century German monk and Protestant Reformers, reformer whose efforts to reform the theology and practice of the Cathol ...
bishop Erik Pontoppidan under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Strix flammea''. The specific epithet is from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
''flammeus'' meaning "flammulated" or "flame-coloured". This owl is now placed with seven other species in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''
Asio ''Asio'' is a genus of typical owls, or true owls, in the family Strigidae. This group has representatives over most of the planet, and the short-eared owl is one of the most widespread of all bird species, breeding in Europe, Asia, North an ...
'' that was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published wo ...
in 1760. Eleven subspecies are recognised: * ''A. f. flammeus'' – (
Pontoppidan Erik Ludvigsen Pontoppidan (24 August 1698 – 20 December 1764) was a Danish author, a Lutheran bishop of the Church of Norway, an historian, and an antiquarian. His Catechism of the Church of Denmark heavily influenced Danish and Norwegian r ...
, 1763)
: the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all spec ...
, found in North America, Europe, North Africa and northern Asia * ''A. f. cubensis'' – Garrido, 2007: found in
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
* ''A. f. domingensis'' – ( Statius Müller, 1776): found on
Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and t ...
* ''A. f. portoricensis'' – Ridgway, 1882: found in
Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...
* ''A. f. bogotensis'' –
Chapman Chapman may refer to: Businesses * Chapman Entertainment, a former British television production company * Chapman Guitars, a guitar company established in 2009 by Rob Chapman * Chapman's, a Canadian ice cream and ice water products manufacturer ...
, 1915
: found in Colombia, Ecuador and northwestern Peru * ''A. f. galapagoensis'' – ( Gould, 1837): found on the
Galápagos Islands The Galápagos Islands ( es, Islas Galápagos) are an archipelago of volcanic islands in the Eastern Pacific, located around the Equator west of the mainland of South America. They form the Galápagos Province of the Republic of Ecuador, with ...
* ''A. f. pallidicaudus'' – Friedmann, 1949: found in
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in ...
, Guyana and
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...
* ''A. f. suinda'' – ( Vieillot, 1817): found from southern Peru and southern Brazil to
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla ...
* ''A. f. sanfordi'' – Bangs, 1919: found on the Falkland Islands * ''A. f. sandwichensis'' – ( A. Bloxam, 1827): Pueo or Hawaiian short-eared owl - found on the
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only ...
an Islands * ''A. f. ponapensis'' – Mayr, 1933: found on eastern Caroline Island


Description

The short-eared owl is a medium-sized owl measuring in length and weighing . It has large eyes, a big head, a short neck, and broad wings. Its bill is short, strong, hooked and black. Its plumage is mottled tawny to brown with a barred tail and wings. The upper breast is significantly streaked. Its flight is characteristically floppy due to its irregular wingbeats. The short-eared owl may also be described as "moth or bat-like" in flight. Wingspans range from . Females are slightly larger than males. The yellow-orange eyes of ''A. flammeus'' are exaggerated by black rings encircling each eye, giving the appearance of them wearing
mascara Mascara is a cosmetic commonly used to enhance the upper and lower eyelashes. It is used to darken, thicken, lengthen, and/or define the eyelashes. Normally in one of three forms—liquid, powder, or cream—the modern mascara product has vari ...
, and large, whitish disks of plumage surrounding the eyes like a mask.


Calls

Short-eared owls have a scratchy bark-like call. Raspy ''waowk, waowk, waowk'' or ''toot-toot-toot-toot-toot'' sounds are common. A loud ''eeee-yerp'' is also heard on breeding grounds. However, short-eared owls are silent on the wintering grounds.


Separation from the long-eared owl

Through much of its range, short-eared owls occurs with the similar-looking long-eared owl. At rest, the ear-tufts of the long-eared owl serve to easily distinguish the two (although long-eared owls can sometimes hold their ear-tufts flat). The iris-colour differs: yellow in the short-eared, and orange in the long-eared, and the black surrounding the eyes is vertical on the long-eared, and horizontal on the short-eared. Overall the short-eared tends to be a paler, sandier bird than the long-eared. There are a number of other ways in which the two species the differ which are best seen when they are flying: a) the short-eared often has a broad white band along the rear edge of the wing, which is not shown by the long-eared; b) on the upperwing, short-eared owls' primary-patches are usually paler and more obvious; c) the band on the upper side of the short-eared owl's tail are usually bolder than those of the long-eared; d) the short-eared's innermost secondaries are often dark-marked, contrasting with the rest of the underwing; e) the long-eared owl has streaking throughout its underparts, whereas on the short-eared the streaking ends at the breast; f) the dark markings on the underside of the tips of the longest primaries are bolder on the short-eared owl; g) the upper parts are coarsely blotched, whereas on the long-eared they are more finely marked. The short-eared owl also differs structurally from the long-eared, having longer, slimmer wings: the long-eared owl has wings shaped more like those of a tawny owl. The long-eared owl generally has different habitat preferences from the short-eared, most often being found concealed in areas with dense wooded thickets. The short-eared owl is often most regularly seen flying about in early morning or late day as it hunts over open habitats.


Distribution and habitat

The short-eared owl occurs on all continents except
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest co ...
and Australia; thus it has one of the most widespread distributions of any bird. ''A. flammeus'' breeds in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
,
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating direction or geography. Etymology The word ''no ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
, the Caribbean,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only ...
and the
Galápagos Islands The Galápagos Islands ( es, Islas Galápagos) are an archipelago of volcanic islands in the Eastern Pacific, located around the Equator west of the mainland of South America. They form the Galápagos Province of the Republic of Ecuador, with ...
. It is partially migratory, moving south in winter from the northern parts of its range. The short-eared owl is known to relocate to areas of higher rodent populations. It will also wander nomadically in search of better food supplies during years when
vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
populations are low. (See
map of the short-eared owl's distribution
across the
New World The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. ...
.)


Behaviour


Breeding

Sexual maturity is attained at one year. Breeding season in the northern hemisphere lasts from March to June, peaking in April. During this time these owls may gather in flocks. During breeding season, the males make great spectacles of themselves in flight to attract females. The male swoops down over the nest flapping its wings in a courtship display. These owls are generally
monogamous Monogamy ( ) is a form of dyadic relationship in which an individual has only one partner during their lifetime. Alternately, only one partner at any one time ( serial monogamy) — as compared to the various forms of non-monogamy (e.g., polyg ...
. The short-eared owl nests on the ground in prairie, tundra, savanna, or meadow habitats. Nests are concealed by low vegetation, and may be lightly lined by weeds, grass, or feathers. Approximately 4 to 7 white eggs are found in a typical
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts ...
, but clutch size can reach up to a dozen eggs in years when
vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
s are abundant. There is one brood per year. The eggs are incubated mostly by the female for 21–37 days. Offspring
fledge Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
at a little over four weeks. This owl is known to lure predators away from its nest by appearing to have a crippled wing.


Food and feeding

Hunting occurs mostly at night, but this owl is known to be diurnal and
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of dayli ...
as well. Its daylight hunting seems to coincide with the high-activity periods of
vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
s, its preferred prey. It tends to fly only feet above the ground in open fields and grasslands until swooping down upon its prey feet-first. Several owls may hunt over the same open area. Its food consists mainly of rodents, especially
vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
s, but it will eat other small mammals such as
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit s ...
s,
mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
,
ground squirrel Ground squirrels are members of the squirrel family of rodents (Sciuridae), which generally live on or in the ground, rather than trees. The term is most often used for the medium-sized ground squirrels, as the larger ones are more commonly known ...
s,
shrew Shrews (family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to diffe ...
s,
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include ''Neotoma'' ( pack rats), ''Bandicota'' (bandicoot ...
s, bats,
muskrat The muskrat (''Ondatra zibethicus'') is a medium-sized semiaquatic rodent native to North America and an introduced species in parts of Europe, Asia, and South America. The muskrat is found in wetlands over a wide range of climates and habita ...
s and moles. It will also occasionally predate smaller birds, especially when near sea-coasts and adjacent wetlands at which time they attack
shorebird 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
s,
tern Terns are seabirds in the family Laridae that have a worldwide distribution and are normally found near the sea, rivers, or wetlands. Terns are treated as a subgroup of the family Laridae which includes gulls and skimmers and consists o ...
s and small
gull Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, ...
s and
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s with semi-regularity. Avian prey is more infrequently preyed on inland and centers on passerines such as
lark Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark oc ...
s,
icterid Icterids () or New World blackbirds make up a family, the Icteridae (), of small to medium-sized, often colorful, New World passerine birds. Most species have black as a predominant plumage color, often enlivened by yellow, orange, or red. The ...
s,
starling Starlings are small to medium-sized passerine birds in the family Sturnidae. The Sturnidae are named for the genus ''Sturnus'', which in turn comes from the Latin word for starling, ''sturnus''. Many Asian species, particularly the larger ones, ...
s,
tyrant flycatcher The tyrant flycatchers (Tyrannidae) are a family of passerine birds which occur throughout North and South America. They are considered the largest family of birds known to exist in the world, with more than 400 species. They are the most diver ...
s and pipits.
Insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...
s supplement the diet and short-eared owls may prey on roaches, grasshoppers,
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s, katydids and
caterpillar Caterpillars ( ) are the larva, larval stage of members of the order Lepidoptera (the insect order comprising butterfly, butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawfly ...
s. Competition can be fierce in North America with the
northern harrier The northern harrier (''Circus hudsonius''), or ring-tailed hawk, is a bird of prey. It breeds throughout the northern parts of the northern hemisphere in Canada and the northernmost USA. The northern harrier migrates to more southerly are ...
, with which the owl shares similar habitat and prey preferences. Both species will readily harass the other when prey is caught. Because of the high pH in the stomach of owls they have a reduced ability to digest bone and other hard parts, so they eject pellets containing the remains of their prey.


Conservation status

The short-eared owl has an estimated global population of 1,200,000 to 2,100,000 and a very large range. On that basis, it is evaluated by the IUCN as a species of
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. Th ...
. It is listed as declining in the southern portion of its United States range. It is common in the northern portion of its breeding range. It is listed as endangered in
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
. Its appearance at the Calverton Executive Airpark on Long Island has prompted the
New York State Department of Environmental Conservation The New York State Department of Environmental Conservation (informally referred to as NYSDEC, DEC, EnCon or NYSENCON) is a department of New York state government. The department guides and regulates the conservation, improvement, and protection ...
to take the lead on ruling whether a massive redevelopment of the airport will receive the necessary environmental permits.


Gallery

File:Asio flammeus -Genovesa Island, Galapagos Islands, Ecuador-8.jpg, up''Asio flammeus galapagoensis'' on the Galapagos Islands, Ecuador File:Short-eared Owl Pangolakha Wildlife Sanctuary East Sikkim India 06.04.2016.jpg, ''Asio flammeus flammeus'' from Pangolakha Wildlife Sanctuary File:Asio flammeus -Fazenda Campo de Ouro, Piraju, Sao Paulo, Brasil-8.jpg, ''Asio flammeus suinda'' in Piraju, São Paulo, Brazil File:Short-eared owl (Asio flammeus) Photograph By Shantanu Kuveskar.jpg, ''Asio flammeus flammeus'' in Mangaon, Maharashtra,
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
File:Starr 090112-0850 Asio flammeus sandwichensis.jpg, ''Asio flammeus sandwichensis'' in Maui File:Sumpfohreule (Asio flammeus) auf Spiekeroog - Nationalpark niedersächsisches Wattenmeer (1).jpg, ''Asio flammeus flammeus'' in Germany


References


Identification

* * * *


External links


Short-eared Owl - ''Asio flammeus''
- USGS Patuxent Bird Identification InfoCenter *
Riverhead, NY ''News Review'' article about the appearance of short-eared owls at EPCAL

Ageing and sexing (PDF; 3.7 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze
* * * * * {{Authority control short-eared owl Holarctic birds Birds of the Americas Birds of North Africa Birds of the Dominican Republic short-eared owl short-eared owl Articles containing video clips Cosmopolitan birds