Shinnery Oak
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''Quercus havardii'' (common names include shinnery oak, shin oak and Havard oak) is a
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
, low-growing, thicket-forming
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
that occupies some in the southern
Great Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, an ...
of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
.
Clones Clone or Clones or Cloning or Cloned or The Clone may refer to: Places * Clones, County Fermanagh * Clones, County Monaghan, a town in Ireland Biology * Clone (B-cell), a lymphocyte clone, the massive presence of which may indicate a pathologi ...
may reach hundreds to thousands of years old, although aboveground stems typically live only 11 to 15 years. Shinnery oak stems are usually tall and codominate the plant community with mid- and tall-grasses, which are usually taller than the oaks. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
honors U.S. Army surgeon and botanist
Valery Havard Valery Havard (February 18, 1846 – November 6, 1927), was a career army officer, physician, author, and botanist. Although he held many notable posts during his military career, he is most well known for his service on the western frontier of th ...
, who contributed much to the knowledge of southwestern plants. The word "shinnery" seems to be derived from ''
chêne Montreux (, , ; frp, Montrolx) is a Swiss municipalities of Switzerland, municipality and List of towns in Switzerland, town on the shoreline of Lake Geneva at the foot of the Swiss Alps, Alps. It belongs to the district of Riviera-Pays-d'Enhaut ...
'' (French for oak), and not from the height of the plant.Oklahoma Biological Survey. ''Quercus havardii Rydb.''
/ref>


Description

Form: A low
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
to or occasionally a small tree, ''Q. havardii'' forms large clonal thickets by extending
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s through the sandy soil where it is usually found. Rhizomes range from in diameter and are concentrated in the upper of soil, although penetration depths of have been reported in shifting dunes. Lateral roots and woody rhizomes are widespread near the soil surface. At least 90% of shinnery oak's biomass is underground, and fortuitous root grafting is common. These underground stems commonly spread to form plants or more in diameter. Single
clones Clone or Clones or Cloning or Cloned or The Clone may refer to: Places * Clones, County Fermanagh * Clones, County Monaghan, a town in Ireland Biology * Clone (B-cell), a lymphocyte clone, the massive presence of which may indicate a pathologi ...
are reported to cover up to and to achieve ages over 13,000 years. Flowers: Shinnery oak is
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...
with both female and male flowers borne separately on the same plant. Male
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged cl ...
s are densely flowered, 1.5–3.8 cm long, and hang downward. Female catkins are long, contain 1 to 5 flowers, and are usually axillary on young shoots. Shinnery oaks are wind pollinated, and flowering occurs in the spring. Acorns:
Acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s develop in one year, maturing in the autumn. Acorns occur alone or in clusters of 2 or 3, and are long by 14–18 mm wide. A scaly cup covers about 1/3 to 1/2 of the nut. On average, acorn crops are produced in 3 out of 10 years. Foliage: The leathery, highly variable leaves are grey green to olive green, have a lustrous upper surface, and are whitish and densely hairy below. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are
alternate Alternative or alternate may refer to: Arts, entertainment and media * Alternative (''Kamen Rider''), a character in the Japanese TV series ''Kamen Rider Ryuki'' * ''The Alternative'' (film), a 1978 Australian television film * ''The Alternative ...
,
simple Simple or SIMPLE may refer to: *Simplicity, the state or quality of being simple Arts and entertainment * ''Simple'' (album), by Andy Yorke, 2008, and its title track * "Simple" (Florida Georgia Line song), 2018 * "Simple", a song by Johnn ...
, with variable
shape A shape or figure is a graphics, graphical representation of an object or its external boundary, outline, or external Surface (mathematics), surface, as opposed to other properties such as color, Surface texture, texture, or material type. A pl ...
(oblong, ovate, or elliptical), and with wavy or shallowly lobed margins. Their length is and width 2 to 4 cm. Twigs: Twigs are brown or grayish, 1–2.5 mm diameter, glabrous or densely covered with short grayish or yellowish hairs, that are lost with age. The buds are dark red-brown, somewhat spherical, about 2 mm long, and sparsely pubescent. Bark: Bark on the larger stems is light gray and scaly.


Taxonomy

Shinnery oak populations in
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
and
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
are considered a variety of shinnery oak ( ''Quercus havardii'' var. ''tuckeri'') by some taxonomists. However,
introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intr ...
of shinnery oak with Gambel oak (''
Quercus gambelii ''Quercus gambelii'', with the common name Gambel oak, is a deciduous small tree or large shrub that is widespread in the foothills and lower mountains of western North America. It is also regionally called scrub oak, oak brush, and white oak. ...
'') and perhaps shrub live oak (''
Quercus turbinella ''Quercus turbinella'' is a North American species of oak known by the common names shrub oak, turbinella oak, shrub live oak, and gray oak. It is native to Arizona, California, New Mexico, Utah, Colorado, and Nevada in the western United States ...
'') make taxonomic identification of these populations difficult. Shinnery oak hybridizes with Mohr oak (''
Quercus mohriana ''Quercus mohriana'', commonly known as the Mohr oak, shin oak or scrub oak, is a North American evergreen shrub or small tree in the white oak group and is native to the south-central United States and north-central Mexico. The species epith ...
'') and post oak (''
Quercus stellata ''Quercus stellata'', the post oak or iron oak, is a North American species of oak in the white oak section. It is a slow-growing oak that lives in dry areas on the edges of fields, tops of ridges also grows in poor soils, and is resistant to ro ...
''). Post oak × shinnery oak hybrids are likely to be a result of post oak's historic range, which extended more westerly than it does today. Mohr oak × shinnery oak hybrids are restricted to habitats intermediate to those occupied by the two species. Mohr oak inhabits limestone soils, and shinnery oak occurs on deep sand soils.


Native range

Shinnery oak is native to western
Oklahoma Oklahoma (; Choctaw language, Choctaw: ; chr, ᎣᎧᎳᎰᎹ, ''Okalahoma'' ) is a U.S. state, state in the South Central United States, South Central region of the United States, bordered by Texas on the south and west, Kansas on the nor ...
, west
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
, eastern
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
, and two isolated populations in southwest
Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the ...
.R.L. McGregor Herbarium. ''Quercus havardii Rydb.'' herbarium records
/ref> A variety also occurs in northeastern Arizona and southeastern Utah.Flora of North America. Quercus havardii: Range Map.
/ref> Sandy plains, sand dunes, and sand hills of the southern Great Plains are typical habitat.


Management

A majority of shinnery oak occurs on private land used for agriculture and/or livestock production. It is considered undesirable on grazing lands, because it competes with better livestock forage and its buds and leaves are toxic to cattle for several weeks in spring. In addition, where cotton is grown near shinnery oak, boll weevils overwinter in the oak litter and infest nearby cotton fields in spring. Thus, most research and other human effort concerning shinnery oak have been devoted to its eradication. The use of chemical and mechanical control methods has been extensive, making it likely that shinnery oak's distribution is decreasing. Most authorities do not recommend complete eradication of shinnery oak, indicating that forage production is typically greater if some shinnery oak remains in the community. Stocking rates and
rotational grazing In agriculture, rotational grazing, as opposed to continuous grazing, describes many systems of pasturing, whereby livestock are moved to portions of the pasture, called paddocks, while the other portions rest. Each paddock must provide all the n ...
patterns exist that reduce the incidence of shinnery oak poisoning of cattle. Excessive control of shinnery oak is controversial because it can open sandy soils to wind erosion and can conflict with wildlife-habitat quality. Shinnery oak sprouts prolifically from rhizomes soon after fire. Researchers described the communities as "extremely fire hardy".


Uses

Because shinnery oak thrives in a harsh environment, it functions as a sand dune stabilizer, protecting sandy soils from wind erosion. It also provides diverse wildlife species cover and food.
Collared peccaries The collared peccary (''Dicotyles tajacu'') is a species of artiodactyl (even-toed) mammal in the family Tayassuidae found in North, Central, and South America. It is the only member of the genus ''Dicotyles''. They are commonly referred to as ...
,
lesser prairie-chicken The lesser prairie chicken (''Tympanuchus pallidicinctus'') is a species in the grouse family. Description It is a medium to large bird, striped white and brown, slightly smaller and paler than its near relative the greater prairie chicken (''T. ...
s,
northern bobwhite The northern bobwhite (''Colinus virginianus''), also known as the Virginia quail or (in its home range) bobwhite quail, is a ground-dwelling bird native to Canada, the United States, Mexico, and Cuba, with introduced populations elsewhere in th ...
s, and many other wildlife species eat shinnery oak acorns.
Deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
,
pronghorn The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American a ...
, and southern plains woodrats browse shinnery oak. Shinnery oak habitats are used by
black-tailed jackrabbit The black-tailed jackrabbit (''Lepus californicus''), also known as the American desert hare, is a common hare of the western United States and Mexico, where it is found at elevations from sea level up to . Reaching a length around , and a ...
s,
desert cottontail The desert cottontail (''Sylvilagus audubonii''), also known as Audubon's cottontail, is a New World cottontail rabbit, and a member of the family Leporidae. Unlike the European rabbit (''Oryctolagus cuniculus''), they do not form social burrow s ...
s,
eastern cottontail The eastern cottontail (''Sylvilagus floridanus'') is a New World cottontail rabbit, a member of the family Leporidae. It is the most common rabbit species in North America. Distribution The eastern cottontail can be found in meadows and shrubby ...
s,
wild turkey The wild turkey (''Meleagris gallopavo'') is an Upland game bird, upland ground bird native to North America, one of two extant species of Turkey (bird), turkey and the heaviest member of the order Galliformes. It is the ancestor to the domestic ...
s, western box turtles, a diversity of arthropods, approximately 25
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
species, and approximately 10
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
species, including the endangered sand dune lizard (''
Sceloporus arenicolus The dunes sagebrush lizard (''Sceloporus arenicolus''), formerly known as the sand dune lizard and the dunes-sagebrush lizard, ''Sceloporus graciosus arenicolus'', a subspecies of sagebrush lizard), is an insectivorous spiny lizard species which ...
''). Many game species also rely on shinnery oak.
Scaled quail The scaled quail (''Callipepla squamata''), also commonly called blue quail or cottontop, is a species of the New World quail family. It is a bluish gray bird found in the arid regions of the Southwestern United States to Central Mexico. This spe ...
and northern bobwhites use shinnery oak habitats extensively, and shinnery oak habitats are considered "the principal home for white-tailed deer in the southern Great Plains". Shinnery oak provides rich habitat for wildlife compared with the surrounding shortgrass plains. Maintaining wildlife habitat and improving vegetation for livestock grazing are often in conflict, and should be carefully considered before undertaking control of shinnery oak.


References


External links

{{Taxonbar, from=Q4112174 havardii Flora of the United States Endemic flora of the United States Trees of the Southwestern United States Trees of the South-Central United States Trees of the North-Central United States Trees of the Southern United States