Sheep or domestic sheep (''Ovis aries'') are
domesticated
Domestication is a sustained multi-generational relationship in which humans assume a significant degree of control over the reproduction and care of another group of organisms to secure a more predictable supply of resources from that group. A ...
,
ruminant
Ruminants (suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
mammals typically kept as
livestock
Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to animal ...
. Although the term ''sheep'' can apply to other species in the genus ''
Ovis
''Ovis'' is a genus of mammals, part of the Caprinae subfamily of the ruminant family Bovidae. Its seven highly sociable species are known as sheep or ovines. Domestic sheep are members of the genus, and are thought to be descended from the w ...
'', in everyday usage it almost always refers to domesticated sheep. Like all ruminants, sheep are members of the
order Artiodactyla
The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
, the
even-toed ungulate
The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
s. Numbering a little over one billion, domestic sheep are also the most numerous species of sheep. An adult female is referred to as a ''ewe'' (), an intact male as a ''ram'', occasionally a ''tup'', a
castrated
Castration is any action, surgical, chemical, or otherwise, by which an individual loses use of the testicles: the male gonad. Surgical castration is bilateral orchiectomy (excision of both testicles), while chemical castration uses pharmac ...
male as a ''wether'', and a young sheep as a ''lamb''.
Sheep are most likely descended from the wild
mouflon
The mouflon (''Ovis gmelini'') is a wild sheep native to Cyprus, the Caspian region from eastern Turkey, Armenia, Azerbaijan, and Iran.
It is thought to be the ancestor of all modern domestic sheep breeds.
Taxonomy
''Ovis gmelini'' was the sc ...
of Europe and Asia, with Iran being a geographic envelope of the domestication center. One of the earliest animals to be domesticated for
agricultural purposes, sheep are raised for
fleeces
Wool is the textile fibre obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have properties similar to animal wool.
As ...
, meat (lamb, hogget or mutton) and
milk
Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digest solid food. Immune factors and immune-modula ...
. A sheep's
wool
Wool is the textile fibre obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have properties similar to animal wool. ...
is the most widely used animal fiber, and is usually harvested by
shearing
Sheep shearing is the process by which the woollen fleece of a sheep is cut off. The person who removes the sheep's wool is called a '' shearer''. Typically each adult sheep is shorn once each year (a sheep may be said to have been "shorn" or ...
. In Commonwealth countries,
ovine
Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticated ...
meat is called
lamb
Lamb or The Lamb may refer to:
* A young sheep
* Lamb and mutton, the meat of sheep
Arts and media Film, television, and theatre
* ''The Lamb'' (1915 film), a silent film starring Douglas Fairbanks Sr. in his screen debut
* ''The Lamb'' (1918 ...
when from younger animals and
mutton
Lamb, hogget, and mutton, generically sheep meat, are the meat of domestic sheep, ''Ovis aries''. A sheep in its first year is a lamb and its meat is also lamb. The meat from sheep in their second year is hogget. Older sheep meat is mutton. Gen ...
when from older ones; in the United States, meat from both older and younger animals is usually called lamb. Sheep continue to be important for wool and meat today, and are also occasionally raised for
pelts, as
dairy animals, or as
model organisms for science.
Sheep husbandry
Sheep farming or sheep husbandry is the raising and breeding of domestic sheep. It is a branch of animal husbandry. Sheep are raised principally for their meat (lamb and mutton), milk ( sheep's milk), and fiber (wool). They also yield sheepski ...
is practised throughout the majority of the inhabited world, and has been fundamental to many civilizations. In the modern era,
Australia,
New Zealand
New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, the southern and central
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
n nations, and the
British Isles
The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
are most closely associated with sheep production.
There is a large
lexicon of unique terms for sheep husbandry which vary considerably by region and
dialect
The term dialect (from Latin , , from the Ancient Greek word , 'discourse', from , 'through' and , 'I speak') can refer to either of two distinctly different types of linguistic phenomena:
One usage refers to a variety of a language that is a ...
. Use of the word ''sheep'' began in
Middle English
Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English ...
as a derivation of the
Old English
Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th c ...
word '; it is both the singular and plural name for the animal. A group of sheep is called a flock. Many other specific terms for the various life stages of sheep exist, generally related to lambing, shearing, and age.
Being a key animal in the history of farming, sheep have a deeply entrenched place in human culture, and find representation in much modern language and
symbology. As livestock, sheep are most often associated with
pastoral,
Arcadian imagery. Sheep figure in many
mythologies
Myth is a folklore genre consisting of narratives that play a fundamental role in a society, such as foundational tales or origin myths. Since "myth" is widely used to imply that a story is not objectively true, the identification of a narrati ...
—such as the
Golden Fleece
In Greek mythology, the Golden Fleece ( el, Χρυσόμαλλον δέρας, ''Chrysómallon déras'') is the fleece of the golden-woolled,, ''Khrusómallos''. winged ram, Chrysomallos, that rescued Phrixus and brought him to Colchis, where ...
—and major religions, especially the
Abrahamic
The Abrahamic religions are a group of religions centered around worship of the God of Abraham. Abraham, a Hebrew patriarch, is extensively mentioned throughout Abrahamic religious scriptures such as the Bible and the Quran.
Jewish tradition ...
traditions. In both ancient and modern religious ritual, sheep are used as
sacrificial animals.
History
The exact line of descent from wild ancestors to domestic sheep is unclear.
The most common hypothesis states that ''Ovis aries'' is descended from the Asiatic (''O. gmelini'') species of
mouflon
The mouflon (''Ovis gmelini'') is a wild sheep native to Cyprus, the Caspian region from eastern Turkey, Armenia, Azerbaijan, and Iran.
It is thought to be the ancestor of all modern domestic sheep breeds.
Taxonomy
''Ovis gmelini'' was the sc ...
; the
European mouflon
The European mouflon (''Ovis aries musimon'') is a feral subspecies of the primitive domestic sheep. It was originally found only on the Mediterranean islands of Corsica and Sardinia, but has since been introduced into many other regions o ...
(''Ovis aries musimon'') is a direct descendant of this population.
Sheep were among the first animals to be domesticated by humankind (although the domestication of dogs probably took place 10 to 20 thousand years earlier); the domestication date is estimated to fall between 11,000 and 9,000 B.C in
Mesopotamia
Mesopotamia ''Mesopotamíā''; ar, بِلَاد ٱلرَّافِدَيْن or ; syc, ܐܪܡ ܢܗܪ̈ܝܢ, or , ) is a historical region of Western Asia situated within the Tigris–Euphrates river system, in the northern part of the ...
[Ensminger, p. 4][Weaver, pp. 11–14][Simmons & Ekarius, p. 2] and possibly around 7,000 B.C. in
Mehrgarh
Mehrgarh (; ur, ) is a Neolithic archaeological site (dated ) situated on the Kacchi Plain of Balochistan in Pakistan. It is located near the Bolan Pass, to the west of the Indus River and between the modern-day Pakistani cities of Quetta, ...
in the Indus Valley. The rearing of sheep for secondary products, and the resulting breed development, began in either southwest Asia or western Europe. Initially, sheep were kept solely for meat, milk and skins. Archaeological evidence from
statuary found at sites in
Iran
Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
suggests that selection for woolly sheep may have begun around 6000 BC,
[Ensminger, p. 5][Weaver, p. 11] and the earliest woven wool garments have been dated to two to three thousand years later.
[Smith et al., p. 8]
Sheep husbandry spread quickly in Europe. Excavations show that in about 6000 BC, during the
Neolithic
The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several p ...
period of prehistory, the Castelnovien people, living around
Châteauneuf-les-Martigues
Châteauneuf-les-Martigues (; oc, Castèunòu dau Martegue) is a commune in the Bouches-du-Rhône department in southern France. The La Mède refinery is nearby and has been in operation since 1935.
Population
See also
* Étang de Berre
* ...
near present-day
Marseille
Marseille ( , , ; also spelled in English as Marseilles; oc, Marselha ) is the prefecture of the French department of Bouches-du-Rhône and capital of the Provence-Alpes-Côte d'Azur region. Situated in the camargue region of southern Fra ...
in the south of France, were among the first in Europe to keep domestic sheep. Practically from its inception,
ancient Greek civilization
Ancient Greece ( el, Ἑλλάς, Hellás) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity ( AD 600), that comprised a loose collection of cultu ...
relied on sheep as primary livestock, and were even said to name individual animals.
[Weaver, p. 13] Ancient Romans
In modern historiography, ancient Rome refers to Roman civilisation from the founding of the city of Rome in the 8th century BC to the collapse of the Western Roman Empire in the 5th century AD. It encompasses the Roman Kingdom (753–509 ...
kept sheep on a wide scale, and were an important agent in the spread of sheep raising.
Pliny the Elder
Gaius Plinius Secundus (AD 23/2479), called Pliny the Elder (), was a Roman author, naturalist and natural philosopher, and naval and army commander of the early Roman Empire, and a friend of the emperor Vespasian. He wrote the encyclopedic ' ...
, in his
Natural History (''Naturalis Historia''), speaks at length about sheep and wool.
European colonists spread the practice to the
New World
The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. ...
from 1493 onwards.
[Ensminger][Weaver, p. 12]
Characteristics
Domestic sheep are relatively small ruminants, usually with a
crimped hair called wool and often with horns forming a
lateral
Lateral is a geometric term of location which may refer to:
Healthcare
*Lateral (anatomy), an anatomical direction
* Lateral cricoarytenoid muscle
* Lateral release (surgery), a surgical procedure on the side of a kneecap
Phonetics
*Lateral co ...
spiral. They differ from their wild relatives and ancestors in several respects, having become uniquely
neotenic
Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compa ...
as a result of selective breeding by humans. A few primitive breeds of sheep retain some of the characteristics of their wild cousins, such as short tails. Depending on breed, domestic sheep may have no horns at all (i.e.
polled), or horns in both sexes, or in males only. Most horned breeds have a single pair, but a few breeds may have several.
Another trait unique to domestic sheep as compared to wild ovines is their wide variation in color. Wild sheep are largely variations of brown hues, and variation within species is extremely limited. Colors of domestic sheep range from pure white to dark chocolate brown, and even spotted or
piebald
A piebald or pied animal is one that has a pattern of unpigmented spots (white) on a pigmented background of hair, feathers or scales. Thus a piebald black and white dog is a black dog with white spots. The animal's skin under the white backgro ...
.
Selection for easily dyeable white fleeces began early in sheep domestication, and as white wool is a
dominant trait
In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and t ...
it spread quickly. However, colored sheep do appear in many modern breeds, and may even appear as a
recessive
In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and t ...
trait in white flocks.
While white wool is desirable for large commercial markets, there is a
niche market for colored fleeces, mostly for
handspinning
Spinning is an ancient textile art in which plant, animal or synthetic fibres are drawn out and twisted together to form yarn. For thousands of years, fibre was spun by hand using simple tools, the spindle and distaff. It was only with the in ...
.
[Weaver] The nature of the fleece varies widely among the breeds, from dense and highly crimped, to long and hairlike. There is variation of wool type and quality even among members of the same flock, so
wool classing
Wool classing is the production of uniform, predictable, low-risk lines of wool, carried out by examining the characteristics of the wool in its raw state and classing (grading) it accordingly. Wool classing is done by a wool classer.
Basis for ...
is a step in the commercial processing of the fibre.
Depending on breed, sheep show a range of heights and weights. Their rate of growth and mature weight is a
heritable
Heredity, also called inheritance or biological inheritance, is the passing on of traits from parents to their offspring; either through asexual reproduction or sexual reproduction, the offspring cells or organisms acquire the genetic informa ...
trait that is often selected for in breeding.
Ewes typically weigh between , and rams between . When all deciduous teeth have erupted, the sheep has 20 teeth. Mature sheep have 32 teeth. As with other ruminants, the front teeth in the lower jaw bite against a hard, toothless pad in the upper jaw. These are used to pick off vegetation, then the rear teeth grind it before it is swallowed. There are eight lower front teeth in ruminants, but there is some disagreement as to whether these are eight
incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s, or six incisors and two incisor-shaped
canines
Canine may refer to:
Zoology and anatomy
* a dog-like Canid animal in the subfamily Caninae
** ''Canis'', a genus including dogs, wolves, coyotes, and jackals
** Dog, the domestic dog
* Canine tooth, in mammalian oral anatomy
People with the surn ...
. This means that the
dental formula
Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolog ...
for sheep is either or There is a large
diastema
A diastema (plural diastemata, from Greek διάστημα, space) is a space or gap between two teeth. Many species of mammals have diastemata as a normal feature, most commonly between the incisors and molars. More colloquially, the condition ...
between the
incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s and the
molars
The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
.
In the first few years of life one can calculate the age of sheep from their front teeth, as a pair of
milk teeth is replaced by larger adult teeth each year, the full set of eight adult front teeth being complete at about four years of age. The front teeth are then gradually lost as sheep age, making it harder for them to feed and hindering the health and productivity of the animal. For this reason, domestic sheep on normal
pasture
Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, cattle, sheep, or sw ...
begin to slowly decline from four years on, and the life expectancy of a sheep is 10 to 12 years, though some sheep may live as long as 20 years.
[Smith et al.] Sheep have good hearing, and are sensitive to noise when being handled.
[Smith et al., p. 5.] Sheep have horizontal slit-shaped pupils, with excellent
peripheral vision
Peripheral vision, or ''indirect vision'', is vision as it occurs outside the point of fixation, i.e. away from the center of gaze or, when viewed at large angles, in (or out of) the "corner of one's eye". The vast majority of the area in th ...
; with visual fields of about 270° to 320°, sheep can see behind themselves without turning their heads.
Many breeds have only short hair on the face, and some have facial wool (if any) confined to the poll and or the area of the mandibular angle; the wide angles of peripheral vision apply to these breeds. A few breeds tend to have considerable wool on the face; for some individuals of these breeds, peripheral vision may be greatly reduced by "wool blindness", unless recently shorn about the face. Sheep have poor
depth perception; shadows and dips in the ground may cause sheep to baulk. In general, sheep have a tendency to move out of the dark and into well-lit areas,
and prefer to move uphill when disturbed. Sheep also have an excellent sense of smell, and, like all species of their genus, have
scent gland
Scent gland are exocrine glands found in most mammals. They produce semi-viscous secretions which contain pheromones and other semiochemical compounds. These odor-messengers indicate information such as status, territorial marking, mood, and s ...
s just in front of the eyes, and interdigitally on the feet. The purpose of these glands is uncertain,
but those on the face may be used in breeding behaviors.
The foot glands might also be related to reproduction,
but alternative functions, such as secretion of a waste product or a scent marker to help lost sheep find their flock, have also been proposed.
[Smith et al., p. 4.]
Comparison with goats
Sheep and
goat
The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s are closely related: both are in the subfamily
Caprinae
The subfamily Caprinae, also sometimes referred to as the tribe Caprini, is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine, or, more informally, a goat-antelope (a ...
. However, they are separate species, so
hybrids rarely occur and are always infertile. A hybrid of a ewe and a buck (a male goat) is called a
sheep-goat hybrid, known as ''geep''. Visual differences between sheep and goats include the beard of goats and divided upper lip of sheep. Sheep tails also hang down, even when short or
docked, while the short tails of goats are held upwards. Also, sheep breeds are often naturally
polled (either in both sexes or just in the female), while naturally polled goats are rare (though many are polled artificially). Males of the two species differ in that buck goats acquire a unique and strong odor during the
rut, whereas rams do not.
Breeds
The domestic sheep is a multi-purpose animal, and the more than 200
breeds
A breed is a specific group of domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist several slight ...
now in existence were created to serve these diverse purposes.
Some sources give a count of a thousand or more breeds,
but these numbers cannot be verified, according to some sources.
However, several hundred breeds of sheep have been identified by the
Food and Agriculture Organization of the UN (FAO), with the estimated number varying somewhat from time to time: e.g. 863 breeds as of 1993, 1314 breeds as of 1995 and 1229 breeds as of 2006.
[FAO. 2007. State of the world's animal genetic resources for food and agriculture] (These numbers exclude extinct breeds, which are also tallied by the FAO.) For the purpose of such tallies, the FAO definition of a breed is "either a subspecific group of domestic livestock with definable and identifiable external characteristics that enable it to be separated by visual appraisal from other similarly defined groups within the same species or a group for which geographical and/or cultural separation from phenotypically similar groups has led to acceptance of its separate identity."
Almost all sheep are classified as being best suited to furnishing a certain product: wool, meat, milk, hides, or a combination in a dual-purpose breed. Other features used when classifying sheep include face color (generally white or black), tail length, presence or lack of horns, and the
topography
Topography is the study of the forms and features of land surfaces. The topography of an area may refer to the land forms and features themselves, or a description or depiction in maps.
Topography is a field of geoscience and planetary sc ...
for which the breed has been developed. This last point is especially stressed in the UK, where breeds are described as either upland (hill or mountain) or lowland breeds.
A sheep may also be of a
fat-tailed type, which is a dual-purpose sheep common in Africa and Asia with larger deposits of fat within and around its tail.
Breeds are often categorized by the type of their wool. Fine wool breeds are those that have wool of great crimp and density, which are preferred for textiles. Most of these were derived from
Merino
The Merino is a breed or group of breeds of domestic sheep, characterised by very fine soft wool. It was established in Spain near the end of the Middle Ages, and was for several centuries kept as a strict Spanish monopoly; exports of the bree ...
sheep, and the breed continues to dominate the world sheep industry. Downs breeds have wool between the extremes, and are typically fast-growing meat and ram breeds with dark faces.
[D’Arcy, J.B., Sheep Management & Wool Technology, NSW University Press, 1986, ] Some major medium wool breeds, such as the
Corriedale, are dual-purpose crosses of long and fine-wooled breeds and were created for high-production commercial flocks. Long wool breeds are the largest of sheep, with long wool and a slow rate of growth. Long wool sheep are most valued for crossbreeding to improve the attributes of other sheep types. For example: the American
Columbia breed was developed by crossing
Lincoln
Lincoln most commonly refers to:
* Abraham Lincoln (1809–1865), the sixteenth president of the United States
* Lincoln, England, cathedral city and county town of Lincolnshire, England
* Lincoln, Nebraska, the capital of Nebraska, U.S.
* Lincol ...
rams (a long wool breed) with fine-wooled
Rambouillet ewes.
Coarse or
carpet
A carpet is a textile floor covering typically consisting of an upper layer of pile attached to a backing. The pile was traditionally made from wool, but since the 20th century synthetic fibers such as polypropylene, nylon, or polyester hav ...
wool sheep are those with a medium to long length wool of characteristic coarseness. Breeds traditionally used for carpet wool show great variability, but the chief requirement is a wool that will not break down under heavy use (as would that of the finer breeds). As the demand for carpet-quality wool declines, some breeders of this type of sheep are attempting to use a few of these traditional breeds for alternative purposes. Others have always been primarily meat-class sheep.
[Wooster]
A minor class of sheep are the
dairy breeds. Dual-purpose breeds that may primarily be meat or wool sheep are often used secondarily as milking animals, but there are a few breeds that are predominantly used for milking. These sheep produce a higher quantity of milk and have slightly longer lactation curves. In the quality of their milk, the fat and protein content percentages of dairy sheep vary from non-dairy breeds, but lactose content does not.
[Pulina et al. p. 2.]
A last group of sheep breeds is that of fur or hair sheep, which do not grow wool at all. Hair sheep are similar to the early domesticated sheep kept before woolly breeds were developed, and are raised for meat and pelts. Some modern breeds of hair sheep, such as the
Dorper
The Dorper is a South African breed of domestic sheep developed by crossing Dorset Horn and the Blackhead Persian sheep. The breed was created through the efforts of the South African Department of Agriculture to breed a meat sheep suitable to ...
, result from crosses between wool and hair breeds. For meat and hide producers, hair sheep are cheaper to keep, as they do not need shearing.
Hair sheep are also more resistant to parasites and hot weather.
With the modern rise of corporate
agribusiness and the decline of localized
family farms, many breeds of sheep are in danger of extinction. The
Rare Breeds Survival Trust
The Rare Breeds Survival Trust is a conservation charity whose purpose is to secure the continued existence and viability of the native farm animal genetic resources (FAnGR) of the United Kingdom. It was founded in 1973 by Joe Henson to pres ...
of the UK lists 22 native breeds as having only 3,000 registered animals (each), and
The Livestock Conservancy
The Livestock Conservancy, formerly known as the American Livestock Breeds Conservancy (ALBC) and prior to that, the American Minor Breeds Conservancy, is a nonprofit organization focused on preserving and promoting rare breeds, also known as "h ...
lists 14 as either "critical" or "threatened". Preferences for breeds with uniform characteristics and fast growth have pushed heritage (or heirloom) breeds to the margins of the sheep industry.
Those that remain are maintained through the efforts of conservation organizations, breed registries, and individual farmers dedicated to their preservation.
Diet
Sheep are
herbivorous
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
mammals. Most breeds prefer to graze on
grass
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
and other short
roughage
Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
, avoiding the taller woody parts of plants that goats readily consume.
[Pugh, pp. 19.] Both sheep and goats use their
lip
The lips are the visible body part at the mouth of many animals, including humans. Lips are soft, movable, and serve as the opening for food intake and in the articulation of sound and speech. Human lips are a tactile sensory organ, and can be ...
s and
tongue
The tongue is a muscular organ in the mouth of a typical tetrapod. It manipulates food for mastication and swallowing as part of the digestive process, and is the primary organ of taste. The tongue's upper surface (dorsum) is covered by taste ...
s to select parts of the plant that are easier to digest or higher in nutrition.
Sheep, however, graze well in
monoculture
In agriculture, monoculture is the practice of growing one crop species in a field at a time. Monoculture is widely used in intensive farming and in organic farming: both a 1,000-hectare/acre cornfield and a 10-ha/acre field of organic kale are ...
pastures where most goats fare poorly.
Like all ruminants, sheep have a complex
digestive system composed of four chambers, allowing them to break down
cellulose
Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell w ...
from stems, leaves, and seed hulls into simpler
carbohydrate
In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or m ...
s. When sheep
graze, vegetation is chewed into a mass called a
bolus, which is then passed into the
rumen
The rumen, also known as a paunch, is the largest stomach compartment in ruminants and the larger part of the reticulorumen, which is the first chamber in the alimentary canal of ruminant animals. The rumen's microbial favoring environment al ...
, via the
reticulum
Reticulum is a small, faint constellation in the southern sky. Its name is Latin for a small net, or reticle—a net of crosshairs at the focus of a telescope eyepiece that is used to measure star positions. The constellation is best viewed b ...
. The rumen is a 19- to 38-liter (5 to 10 gallon) organ in which feed is
fermented
Fermentation is a metabolic process that produces chemical changes in organic substrates through the action of enzymes. In biochemistry, it is narrowly defined as the extraction of energy from carbohydrates in the absence of oxygen. In food p ...
.
[Simmons & Ekarius, p. 146.] The fermenting organisms include bacteria, fungi, and protozoa. (Other important rumen organisms include some archaea, which produce methane from carbon dioxide.) The bolus is periodically regurgitated back to the mouth as
cud
Cud is a portion of food that returns from a ruminant's stomach to the mouth to be chewed for the second time. More precisely, it is a bolus of semi-degraded food regurgitated from the reticulorumen of a ruminant. Cud is produced during the phy ...
for additional chewing and
salivation.
After fermentation in the rumen, feed passes into the
reticulum
Reticulum is a small, faint constellation in the southern sky. Its name is Latin for a small net, or reticle—a net of crosshairs at the focus of a telescope eyepiece that is used to measure star positions. The constellation is best viewed b ...
and the
omasum
The omasum, also known as the bible, the fardel, the manyplies and the psalterium, is the third compartment of the stomach in ruminants. The omasum comes after the rumen and reticulum and before the abomasum. Different ruminants have different oma ...
; special feeds such as grains may bypass the rumen altogether. After the first three chambers, food moves into the
abomasum
The abomasum, also known as the maw,[The Cham ...](_blank)
for final digestion before processing by the
intestines. The abomasum is the only one of the four chambers analogous to the human stomach, and is sometimes called the "true stomach".
Other than forage, the other staple feed for sheep is
hay
Hay is grass, legumes, or other herbaceous plants that have been cut and dried to be stored for use as animal fodder, either for large grazing animals raised as livestock, such as cattle, horses, goats, and sheep, or for smaller domesticat ...
, often during the winter months. The ability to thrive solely on pasture (even without hay) varies with breed, but all sheep can survive on this diet.
Also included in some sheep's diets are
mineral
In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid chemical compound with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. (2 ...
s, either in a trace mix or in
licks. Feed provided to sheep must be specially formulated, as most cattle, poultry, pig, and even some goat feeds contain levels of
copper
Copper is a chemical element with the symbol Cu (from la, cuprum) and atomic number 29. It is a soft, malleable, and ductile metal with very high thermal and electrical conductivity. A freshly exposed surface of pure copper has a pinkis ...
that are lethal to sheep.
The same danger applies to mineral supplements such as salt licks.
Grazing behavior
Sheep follow a diurnality, diurnal pattern of activity, feeding from dawn to dusk, stopping sporadically to rest and chew their
cud
Cud is a portion of food that returns from a ruminant's stomach to the mouth to be chewed for the second time. More precisely, it is a bolus of semi-degraded food regurgitated from the reticulorumen of a ruminant. Cud is produced during the phy ...
. Ideal pasture for sheep is not lawnlike grass, but an array of
grass
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
es, legumes and forbs. Types of land where sheep are raised vary widely, from pastures that are seeded and improved intentionally to rough, native lands. Common plants toxic to sheep are present in most of the world, and include (but are not limited to) cherry, some oaks and acorns, tomato, Taxus, yew, rhubarb, potato, and rhododendron.
Effects on pasture
Sheep are largely grazing herbivores, unlike Browsing (predation), browsing animals such as goats and deer that prefer taller foliage. With a much narrower face, sheep crop plants very close to the ground and can overgraze a pasture much faster than cattle.
For this reason, many shepherds use managed intensive rotational grazing, where a flock is rotated through multiple pastures, giving plants time to recover.
Paradoxically, sheep can both cause and solve the spread of invasive species, invasive plant species. By disturbing the natural state of pasture, sheep and other livestock can pave the way for invasive plants. However, sheep also prefer to eat invasives such as cheatgrass, Euphorbia virgata, leafy spurge, kudzu and spotted knapweed over native species such as sagebrush, making grazing sheep effective for conservation grazing. Research conducted in Imperial County, California compared lamb grazing with herbicides for weed control in seedling alfalfa fields. Three trials demonstrated that grazing lambs were just as effective as herbicides in controlling winter weeds. Entomologists also compared grazing lambs to insecticides for insect control in winter alfalfa. In this trial, lambs provided insect control as effectively as insecticides.
Behavior
Flock behavior
Sheep are Herd animal, flock animals and strongly gregarious; much sheep behavior can be understood on the basis of these tendencies. The dominance hierarchy of sheep and their natural inclination to follow a leader to new pastures were the pivotal factors in sheep being one of the first domesticated livestock species.
[Budiansky] Furthermore, in contrast to the red deer and gazelle (two other ungulates of primary importance to meat production in prehistoric times), sheep do not defend Territory (animal), territories although they do form home ranges.
[Clutton-Brock, J., (1987). A Natural History of Domesticated Mammals. Cambridge University Press, Cambridge pp.55] All sheep have a tendency to congregate close to other members of a flock, although this behavior varies with breed,
[ and sheep can become stressed when separated from their flock members.] During flocking, sheep have a strong tendency to follow, and a leader may simply be the first individual to move. Relationships in flocks tend to be closest among related sheep: in mixed-breed flocks, subgroups of the same breed tend to form, and a ewe and her direct descendants often move as a unit within large flocks. Sheep can become hefted to one particular local pasture (heft) so they do not roam freely in unfenced landscapes. Lambs learn the heft from ewes and if whole flocks are culled it must be retaught to the replacement animals.
Flock behaviour in sheep is generally only exhibited in groups of four or more sheep; fewer sheep may not react as expected when alone or with few other sheep. Being a prey species, the primary defense mechanism of sheep is to flee from danger when their flight zone is entered. Cornered sheep may charge and butt, or threaten by hoof stamping and adopting an aggressive posture. This is particularly true for ewes with newborn lambs.
In regions where sheep have no natural predators, none of the native breeds of sheep exhibit a strong flocking behavior.
Herding
Farmers exploit flocking behavior to keep sheep together on unfenced pastures such as hill farming, and to move them more easily. For this purpose shepherds may use herding dogs in this effort, with a highly bred herding ability. Sheep are food-oriented, and association of humans with regular feeding often results in sheep soliciting people for food. Those who are moving sheep may exploit this behavior by leading sheep with buckets of feed.
Dominance hierarchy
Sheep establish a Dominance (ethology), dominance hierarchy through fighting, threats and competitiveness. Dominant animals are inclined to be more aggressive with other sheep, and usually feed first at wikt:trough, troughs. Primarily among rams, horn size is a factor in the flock hierarchy.[Budiansky p. 78.] Rams with different size horns may be less inclined to fight to establish the dominance order, while rams with similarly sized horns are more so. Merinos have an almost linear hierarchy whereas there is a less rigid structure in Border Leicesters when a competitive feeding situation arises.
In sheep, position in a moving flock is highly correlated with social dominance, but there is no definitive study to show consistent voluntary leadership by an individual sheep.
Intelligence and learning ability
Sheep are frequently thought of as Animal cognition, unintelligent animals. Their flocking behavior and quickness to flee and panic can make shepherding a difficult endeavor for the uninitiated. Despite these perceptions, a University of Illinois system, University of Illinois monograph on sheep reported their intelligence to be just below that of pigs and on par with that of cattle.
Sheep can recognize individual human and ovine faces and remember them for years; they can remember 50 other different sheep faces for over two years; they can recognize and are attracted to individual sheep and humans by their faces, as they possess similar specialized neural systems in the temporal lobe, temporal and frontal lobes of their brains to humans and have a greater involvement of the right brain hemisphere. In addition to long-term facial recognition of individuals, sheep can also differentiate emotional states through facial characteristics. If worked with patiently, sheep may learn their names, and many sheep are trained to be led by halter for showing and other purposes. Sheep have also responded well to clicker training. Sheep have been used as pack animals; Tibetan people, Tibetan nomads distribute baggage equally throughout a flock as it is herded between living sites.
It has been reported that some sheep have apparently shown problem-solving abilities; a flock in West Yorkshire, England allegedly found a way to get over cattle grids by rolling on their backs, although documentation of this has relied on anecdotal accounts.
Vocalisations
Sounds made by domestic sheep include bleats, grunts, rumbles and snorts. Bleating ("baaing") is used mostly for contact communication, especially between dam and lambs, but also at times between other flock members.[Lynch, J.J., G. N Hinch and D. B. Adams. (1992). ''The Behaviour Of Sheep: Biological Principles And Implications For Production.'' CABI, Wallingford. 237 pp.] The bleats of individual sheep are distinctive, enabling the ewe and her lambs to recognize each other's vocalizations. Vocal communication between lambs and their dam declines to a very low level within several weeks after parturition.[ A variety of bleats may be heard, depending on sheep age and circumstances. Apart from contact communication, bleating may signal distress, frustration or impatience; however, sheep are usually silent when in pain. Isolation commonly prompts bleating by sheep.][Dwyer, C.M. (ed.) (2008). ''The Welfare Of Sheep.'' CABI, Wallingford, UK. 366 pp.] Pregnant ewes may grunt when in labor. Rumbling sounds are made by the ram during courting; somewhat similar rumbling sounds may be made by the ewe,[ especially when with her neonate lambs. A snort (explosive exhalation through the nostrils) may signal aggression or a warning,][ and is often elicited from startled sheep.
]
Senses
In sheep breeds lacking facial wool, the visual field is wide. In 10 sheep (Cambridge, Lleyn and Welsh Mountain breeds, which lack facial wool), the visual field ranged from 298° to 325°, averaging 313.1°, with binocular overlap ranging from 44.5° to 74°, averaging 61.7°.[Piggins, D. and C. J. C. Phillips. 1996. The eye of the domesticated sheep and its implications for vision. Animal Science. 62: 301–308.] In some breeds, unshorn facial wool can limit the visual field; in some individuals, this may be enough to cause "wool blindness". In 60 Merinos, visual fields ranged from 219.1° to 303.0°, averaging 269.9°, and the binocular field ranged from 8.9° to 77.7°, averaging 47.5°; 36% of the measurements were limited by wool,[Hutson, G. D. 1980. Visual field, restricted vision and sheep movement in laneways. Appl. Anim. Ethol. 6: 175–187.] although photographs of the experiments indicate that only limited facial wool regrowth had occurred since shearing. In addition to facial wool (in some breeds), visual field limitations can include ears and (in some breeds) horns,[ so the visual field can be extended by tilting the head. Sheep eyes exhibit very low hyperopia and little Astigmatism (eye), astigmatism. Such visual characteristics are likely to produce a well-focused retinal image of objects in both the middle and long distance.][ Because sheep eyes have no accommodation (eye), accommodation, one might expect the image of very near objects to be blurred, but a rather clear near image could be provided by the Tapetum lucidum, tapetum and large retinal image of the sheep's eye, and adequate close vision may occur at muzzle length.][ Good depth perception, inferred from the sheep's sure-footedness, was confirmed in "visual cliff" experiments;][ behavioral responses indicating depth perception are seen in lambs at one day old. Sheep are thought to have colour vision, and can distinguish between a variety of colours: black, red, brown, green, yellow and white.][Alexander, G. and Shillito, E.E. (1978). Maternal responses in Merino ewes to artificially coloured lambs. Applied Animal Ethology, 4: 141–152] Sight is a vital part of sheep communication, and when grazing, they maintain visual contact with each other.[Kilgour, R., (1977). Design sheep yards to suit the whims of sheep. N.Z. Farmer, 98(6): 29–31] Each sheep lifts its head upwards to check the position of other sheep in the flock. This constant monitoring is probably what keeps the sheep in a flock as they move along grazing. Sheep become stressed when isolated; this stress is reduced if they are provided with a mirror, indicating that the sight of other sheep reduces stress.[Parrott, R.F., (1990). Physiological responses to isolation in sheep. Social Stress in Domestic Animals, Kluwer Academic Publishers, Dordrecht, Netherlands: 1990. 212 -226]
Taste is the most important sense in sheep, establishing forage preferences, with sweet and sour plants being preferred and bitter plants being more commonly rejected. Touch and sight are also important in relation to specific plant characteristics, such as succulence and growth form.[Krueger, W.C., Laycock, W.A. and Price, D.A., (1974). Relationships of taste, smell, sight and touch on forage selection. Journal of Range Management, 27(4): 258–262]
The ram uses his vomeronasal organ (sometimes called the Jacobson's organ) to sense the pheromones of ewes and detect when they are in estrus. The ewe uses her vomeronasal organ for early recognition of her neonate lamb.
Reproduction
Sheep follow a similar Reproduction, reproductive strategy to other herd animals. A group of ewes is generally mated by a single ram, who has either been chosen by a breeder or (in feral populations) has established dominance through physical contest with other rams. Most sheep are seasonal breeders, although some are able to breed year-round. Ewes generally reach sexual maturity at six to eight months old, and rams generally at four to six months. However, there are exceptions. For example, Finnsheep ewe lambs may reach puberty as early as 3 to 4 months, and Merino ewes sometimes reach puberty at 18 to 20 months. Ewes have estrus cycles about every 17 days, during which they emit a scent and indicate readiness through physical displays towards rams. A minority of rams (8% on average) display a preference for Homosexual behavior in animals, homosexuality and a small number of the females that were accompanied by a male fetus ''in utero'' are freemartins (female animals that are behaviorally masculine and lack functioning Ovary, ovaries).
In feral sheep, rams may fight during the rut to determine which individuals may mate with ewes. Rams, especially unfamiliar ones, will also fight outside the breeding period to establish dominance; rams can kill one another if allowed to mix freely. During the rut, even usually friendly rams may become aggressive towards humans due to increases in their hormone levels.
After mating, sheep have a gestation period of about five months, and normal labor takes one to three hours. Although some breeds regularly throw larger litters of lambs, most produce single or twin lambs. During or soon after labor, ewes and lambs may be confined to small Glossary of sheep husbandry, lambing jugs, small pens designed to aid both careful observation of ewes and to cement the bond between them and their lambs.
Ovine obstetrics can be problematic. By selectively breeding ewes that produce multiple offspring with higher birth weights for generations, sheep producers have inadvertently caused some domestic sheep to have difficulty lambing; balancing ease of lambing with high productivity is one of the dilemmas of sheep breeding. In the case of any such problems, those present at lambing may assist the ewe by extracting or repositioning lambs. After the birth, ewes ideally break the amniotic sac (if it is not broken during labor), and begin licking clean the lamb. Most lambs will begin standing within an hour of birth. In normal situations, lambs nurse after standing, receiving vital colostrum milk. Lambs that either fail to nurse or are rejected by the ewe require help to survive, such as bottle-feeding or fostering by another ewe.
Most lambs begin life being born outdoors. After lambs are several weeks old, Glossary of sheep husbandry, lamb marking (Glossary of sheep husbandry, ear tagging, Docking (animal), docking, mulesing, and castrating) is carried out.[Wooster] Vaccinations are usually carried out at this point as well. Ear tags with numbers are attached, or ear marks are applied, for ease of later identification of sheep. Docking and castration are commonly done after 24 hours (to avoid interference with maternal bonding and consumption of colostrum) and are often done not later than one week after birth, to minimize pain, stress, recovery time and complications. The first course of vaccinations (commonly anti-clostridial) is commonly given at an age of about 10 to 12 weeks; i.e. when the concentration of maternal antibodies passively acquired via colostrum is expected to have fallen low enough to permit development of active immunity. Ewes are often revaccinated annually about 3 weeks before lambing, to provide high antibody concentrations in colostrum during the first several hours after lambing. Ram lambs that will either be slaughtered or separated from ewes before sexual maturity are not usually castrated. Objections to all these procedures have been raised by animal rights groups, but farmers defend them by saying they save money, and inflict only temporary pain.[Simmons & Ekarius]
Health
Sheep may fall victim to poisons, List of infectious sheep and goat diseases, infectious diseases, and physical injuries. As a prey species, a sheep's system is adapted to hide the obvious signs of illness, to prevent being targeted by predators. However, some signs of ill health are obvious, with sick sheep eating little, vocalizing excessively, and being generally listless. Throughout history, much of the money and labor of sheep husbandry has aimed to prevent sheep ailments. Historically, shepherds often created remedies by experimentation on the farm. In some developed countries, including the United States, sheep lack the economic importance for drug companies to perform expensive clinical trials required to approve more than a relatively limited number of drugs for ovine use. However, extra-label drug use in sheep production is permitted in many jurisdictions, subject to certain restrictions. In the US, for example, regulations governing extra-label drug use in animals are found in 21 CFR (Code of Federal Regulations) Part 530. In the 20th and 21st centuries, a minority of sheep owners have turned to alternative treatments such as homeopathy, herbalism and even traditional Chinese medicine to treat sheep veterinary problems. Despite some favorable anecdotal evidence, the effectiveness of alternative veterinary medicine has been met with skepticism in scientific journals. The need for traditional anti-parasite drugs and antibiotics is widespread, and is the main impediment to certified organic farming with sheep.
Many breeders take a variety of Preventive medicine, preventive measures to ward off problems. The first is to ensure all sheep are healthy when purchased. Many buyers avoid outlets known to be clearing houses for animals culled from healthy flocks as either sick or simply inferior. This can also mean maintaining a closed flock, and Quarantine, quarantining new sheep for a month. Two fundamental preventive programs are maintaining good nutrition and reducing stress in the sheep. Restraint, isolation, loud noises, novel situations, pain, heat, extreme cold, fatigue and other stressors can lead to secretion of cortisol, a stress hormone, in amounts that may indicate welfare problems.[Gregory, N. G. 1998. Animal welfare and meat science. CABI, Wallingford, UK. 298 pp.][Moberg, G. P. and J. A. Mench. 2000. The biology of animal stress: basic principles and implications for welfare. CABI, Wallingford, UK. pp. 1–21.] Excessive stress can compromise the immune system. "Shipping fever" (pneumonic mannheimiosis, formerly called pasteurellosis) is a disease of particular concern, that can occur as a result of stress, notably during transport and (or) handling. Pain, fear and several other stressors can cause secretion of epinephrine (adrenaline). Considerable epinephrine secretion in the final days before slaughter can adversely affect meat quality (by causing glycogenolysis, removing the substrate for normal post-slaughter acidification of meat) and result in meat becoming more susceptible to colonization by spoilage bacteria. Because of such issues, low-stress handling is essential in sheep management. Avoiding poisoning is also important; common poisons are pesticide sprays, inorganic fertilizer, motor oil, as well as radiator coolant containing ethylene glycol.
Common forms of preventive medication for sheep are vaccinations and treatments for parasites. Both external and internal parasites are the most prevalent malady in sheep, and are either fatal, or reduce the productivity of flocks. Worms are the most common internal parasites. They are ingested during grazing, incubate within the sheep, and are expelled through the digestive system (beginning the cycle again). Oral anti-parasitic medicines, known as Deworming, drenches, are given to a flock to treat worms, sometimes after worm eggs in the feces has been counted to assess infestation levels. Afterwards, sheep may be moved to a new pasture to avoid ingesting the same parasites. External sheep parasites include: lice (for different parts of the body), sheep Hippoboscidae, keds, nose bots, Psorergates ovis, sheep itch mites, and maggots. Keds are blood-sucking parasites that cause general malnutrition and decreased productivity, but are not fatal. Maggots are those of the bot fly and the blow-fly, commonly ''Common green bottle fly, Lucilia sericata'' or its relative ''L. cuprina''. Fly maggots cause the extremely destructive condition of flystrike. Flies lay their eggs in wounds or wet, manure-soiled wool; when the maggots hatch they burrow into a sheep's flesh, eventually causing death if untreated. In addition to other treatments, crutching (shearing wool from a sheep's rump) is a common preventive method. Some countries allow mulesing, a practice that involves stripping away the skin on the rump to prevent fly-strike, normally performed when the sheep is a lamb. Nose bots are fly larvae that inhabit a sheep's Paranasal sinus, sinuses, causing breathing difficulties and discomfort. Common signs are a discharge from the nasal passage, sneezing, and frantic movement such as head shaking. External parasites may be controlled through the use of Glossary of sheep husbandry, backliners, sprays or immersive sheep dips.
A wide array of bacterial and viral diseases affect sheep. Diseases of the hoof, such as foot rot and foot scald may occur, and are treated with footbaths and other remedies. Foot rot is present in over 97% of flocks in the UK. These painful conditions cause lameness and hinder feeding. Ovine Johne's disease is a wasting disease that affects young sheep. Bluetongue disease is an insect-borne illness causing fever and inflammation of the mucous membranes. Ovine rinderpest (or ''peste des petits ruminants'') is a highly contagious and often fatal virus, viral disease affecting sheep and goats. Sheep may also be affected by primary or secondary photosensitization. Tetanus can also afflict sheep through wounds from Sheep shearing, shearing, Docking (animal), docking, castration, or vaccination. The organism also can be introduced into the reproductive tract by unsanitary humans who assist ewes during lambing.
A few sheep conditions are transmissible to humans. Orf (animal disease), Orf (also known as scabby mouth, contagious ecthyma or soremouth) is a skin disease leaving lesions that is transmitted through skin-to-skin contact. Cutaneous anthrax is also called woolsorter's disease, as the spores can be transmitted in unwashed wool. More seriously, the organisms that can cause spontaneous enzootic abortion in sheep are easily transmitted to pregnant women. Also of concern are the prion disease scrapie and the virus that causes foot-and-mouth disease (FMD), as both can devastate flocks. The latter poses a slight risk to humans. During the 2001 FMD pandemic in the UK, hundreds of sheep were culled and some rare British breeds were at risk of extinction due to this.
Of the 600,300 sheep lost to the US economy in 2004, 37.3% were lost to predators, while 26.5% were lost to some form of disease. Poisoning accounted for 1.7% of non-productive deaths.[
]
Predators
Other than parasites and disease, predation is a threat to sheep and the profitability of sheep raising. Sheep have little ability to defend themselves, compared with other species kept as livestock. Even if sheep survive an attack, they may die from their injuries or simply from panic. However, the impact of predation varies dramatically with region. In Africa, Australia, the Americas, and parts of Europe and Asia predators are a serious problem. In the United States, for instance, over one third of sheep deaths in 2004 were caused by predation. In contrast, other nations are virtually devoid of sheep predators, particularly islands known for extensive sheep husbandry. Worldwide, Canidae, canids—including the domestic dog—are responsible for most sheep deaths. Other animals that occasionally prey on sheep include: felines, bears, birds of prey, ravens and Razorback, feral hogs.
Sheep producers have used a wide variety of measures to combat predation. Pre-modern shepherds used their own presence, livestock guardian dogs, and protective structures such as barns and fencing. Fencing (both regular and Electric fence, electric), penning sheep at night and lambing indoors all continue to be widely used. More modern shepherds used guns, Animal trapping, traps, and poisons to kill predators, causing significant decreases in predator populations. In the wake of the environmental and conservation movements, the use of these methods now usually falls under the purview of specially designated government agencies in most developed countries.
The 1970s saw a resurgence in the use of livestock guardian dogs and the development of new methods of predator control by sheep producers, many of them non-lethal. Donkeys and guard llamas have been used since the 1980s in sheep operations, using the same basic principle as livestock guardian dogs. Interspecific pasturing, usually with larger livestock such as cattle or horses, may help to deter predators, even if such species do not actively guard sheep. In addition to animal guardians, contemporary sheep operations may use non-lethal predator deterrents such as motion-activated lights and noisy alarms.
Economic importance
Sheep are an important part of the global agricultural economy. However, their once vital status has been largely replaced by other livestock species, especially the pig, chicken, and cow. People's Republic of China, China, Australia, India, and Iran
Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
have the largest modern flocks, and serve both local and exportation needs for wool and mutton. Other countries such as New Zealand have smaller flocks but retain a large international economic impact due to their export of sheep products. Sheep also play a major role in many local economies, which may be niche markets focused on organic or sustainable agriculture and Locavore, local food customers. Especially in developing country, developing countries, such flocks may be a part of subsistence agriculture rather than a system of trade. Sheep themselves may be a medium of trade in Barter, barter economies.
Domestic sheep provide a wide array of raw materials. Wool was one of the first textiles, although in the late 20th century wool prices began to fall dramatically as the result of the popularity and cheap prices for synthetic fabrics. For many sheep owners, the cost of shearing is greater than the possible profit from the fleece, making subsisting on wool production alone practically impossible without farm subsidies. Fleeces are used as material in making alternative products such as wool insulation. In the 21st century, the sale of meat is the most profitable enterprise in the sheep industry, even though far less sheep meat is consumed than chicken, pork or beef.
Sheepskin is likewise used for making clothes, footwear, rugs, and other products. Byproducts from the slaughter of sheep are also of value: sheep tallow can be used in candle and soap making, sheep bone and cartilage has been used to furnish carved items such as dice and buttons as well as rendered glue and gelatin. Sheep intestine can be formed into sausage casings, and lamb intestine has been formed into surgical sutures, as well as strings for musical instruments and tennis rackets. Sheep droppings, which are high in cellulose
Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell w ...
, have even been sterilized and mixed with traditional Wood pulp, pulp materials to make paper. Of all sheep byproducts, perhaps the most valuable is lanolin: the waterproof, fatty substance found naturally in sheep's wool and used as a base for innumerable cosmetics and other products.
Some farmers who keep sheep also make a profit from live sheep. Providing lambs for youth programs such as 4-H and competition at agricultural shows is often a dependable avenue for the sale of sheep. Farmers may also choose to focus on a particular breed of sheep in order to sell registered purebred animals, as well as provide a ram rental service for breeding. A new option for deriving profit from live sheep is the rental of flocks for grazing; these "Lawn mower, mowing services" are hired in order to keep unwanted vegetation down in public spaces and to lessen fire hazard.
Despite the falling demand and price for sheep products in many markets, sheep have distinct economic advantages when compared with other livestock. They do not require expensive housing, such as that used in the intensive farming of chickens or pigs. They are an efficient use of land; roughly six sheep can be kept on the amount that would suffice for a single cow or horse. Sheep can also consume plants, such as noxious weeds, that most other animals will not touch, and produce more young at a faster rate. Also, in contrast to most livestock species, the cost of raising sheep is not necessarily tied to the price of feed crops such as grain, soybeans and corn. Combined with the lower cost of quality sheep, all these factors combine to equal a lower overhead (business), overhead for sheep producers, thus entailing a higher profitability potential for the small farmer. Sheep are especially beneficial for independent producers, including family farms with limited resources, as the sheep industry is one of the few types of animal agriculture that has not been vertically integrated by agribusiness. However, small flocks, from 10 to 50 ewes, often are not profitable because they tend to be poorly managed. The primary reason is that mechanization is not feasible, so return per hour of labor is not maximized. Small farm flocks generally are used simply to control weeds on irrigation ditches or maintained as a hobby.
As food
Sheep meat and milk were one of the earliest staple proteins consumed by human civilization after the transition from Hunter-gatherer, hunting and gathering to agriculture. Sheep meat prepared for food is known as either mutton or lamb, and approximately 540 million sheep are slaughtered each year for meat worldwide. "Mutton" is derived from the Old French ''moton'', which was the word for sheep used by the Normans, Anglo-Norman rulers of much of the British Isles in the Middle Ages. This became the name for sheep meat in English, while the Old English language, Old English word ''sceap'' was kept for the live animal. Throughout modern history, "mutton" has been limited to the meat of mature sheep usually at least two years of age; "lamb" is used for that of immature sheep less than a year.
In the 21st century, the nations with the highest consumption of sheep meat are the Arab States of the Persian Gulf, New Zealand, Australia, Greece, Uruguay, the United Kingdom and Ireland. These countries eat 14–40 lbs (3–18 kg) of sheep meat per capita, per annum. Sheep meat is also popular in France, Africa (especially the Arab World), the Caribbean, the rest of the Middle East, India, and parts of China. This often reflects a history of sheep production. In these countries in particular, dishes comprising alternative cuts and offal may be popular or traditional. Sheep testicles—called animelles or lamb fries—are considered a delicacy in many parts of the world. Perhaps the most unusual dish of sheep meat is the Scottish haggis, composed of various sheep offal, innards cooked along with oatmeal and chopped onions inside its stomach. In comparison, countries such as the U.S. consume only a pound or less (under 0.5 kg), with Americans eating 50 pounds (22 kg) of pork and 65 pounds (29 kg) of beef. In addition, such countries rarely eat mutton, and may favor the more expensive cuts of lamb: mostly Meat chop, lamb chops and leg of lamb.
Though sheep's milk may be drunk rarely in fresh form, today it is used predominantly in cheese and yogurt making. Sheep have only two teats, and produce a far smaller volume of milk than cows. However, as sheep's milk contains far more fat, Milk solids, solids, and minerals than cow's milk, it is ideal for the cheese-making process. It also resists contamination during cooling better because of its much higher calcium content. Well-known cheeses made from sheep milk include the Feta of Bulgaria and Greece, Roquefort of France, Manchego from Spain, the Pecorino Romano cheese, Pecorino Romano (the Italian language, Italian word for sheep is ''pecore'') and Ricotta of Italy. Yogurts, especially some forms of strained yogurt, may also be made from sheep milk. Many of these products are now often made with cow's milk, especially when produced outside their country of origin. Sheep milk contains 4.8% lactose, which may affect those who are lactose intolerance, intolerant.
As with other domestic animals, the meat of uncastrated males is inferior in quality, especially as they grow. A "bucky" lamb is a lamb which was not castrated early enough, or which was castrated improperly (resulting in one testicle being retained). These lambs are worth less at market.
In science
Sheep are generally too large and reproduce too slowly to make ideal research subjects, and thus are not a common model organism. They have, however, played an influential role in some fields of science. In particular, the Roslin Institute of Edinburgh, Scotland used sheep for genetics research that produced groundbreaking results. In 1995, two ewes named Megan and Morag (cloned sheep), Megan and Morag were the first mammals Cloning, cloned from Cellular differentiation, differentiated cells, also referred to as gynomerogony. A year later, a Finnish Dorset sheep named Dolly (sheep), Dolly, dubbed "the world's most famous sheep" in ''Scientific American'', was the first mammal to be cloned from an adult somatic cell. Following this, Polly and Molly were the first mammals to be simultaneously cloned and transgenic.
As of 2008, the sheep genome has not been fully sequenced, although a detailed genetic linkage, genetic map has been published, and a draft version of the complete genome produced by assembling sheep DNA sequences using information given by the genomes of other mammals. In 2012, a transgenic sheep named "Peng Peng" was cloned by Chinese scientists, who spliced his genes with that of a roundworm (Caenorhabditis elegans, C. elegans) in order to increase production of fats healthier for human consumption.
In the study of natural selection, the population of Soay sheep that remain on the island of Hirta have been used to explore the relation of body size and coloration to reproductive success. Soay sheep come in several colors, and researchers investigated why the larger, darker sheep were in decline; this occurrence contradicted the rule of thumb that larger members of a population tend to be more successful reproductively. The feral Soays on Hirta are especially useful subjects because they are isolated.
Sheep are one of the few animals where the molecular basis of the diversity of male sexual preferences has been examined. However, this research has been controversial, and much publicity has been produced by a study at the Oregon Health and Science University that investigated the mechanisms that produce homosexuality in rams. Organizations such as PETA campaigned against the study, accusing scientists of trying to cure homosexuality in the sheep. OHSU and the involved scientists vehemently denied such accusations.
Domestic sheep are sometimes used in medical research, particularly for researching cardiovascular physiology, in areas such as hypertension and heart failure. Pregnant sheep are also a useful model for human pregnancy, and have been used to investigate the effects on fetal development of malnutrition and Hypoxia (medical), hypoxia. In behavioral sciences, sheep have been used in isolated cases for the study of Face perception, facial recognition, as their mental process of recognition is qualitatively similar to humans.
Cultural impact
Sheep have had a strong presence in many cultures, especially in areas where they form the most common type of livestock. In the English language, to call someone a sheep or ovine may allude that they are timid and easily led. In contradiction to this image, male sheep are often used as symbols of virility and power; the logos of the Los Angeles Rams football team and the Dodge Ram pickup truck allude to males of the bighorn sheep, ''Ovis canadensis''.
Counting sheep is popularly said to be an aid to sleep, and some Yan Tan Tethera, ancient systems of counting sheep persist today. Sheep also enter in colloquial sayings and idiom frequently with such phrases as "black sheep". To call an individual a black sheep implies that they are an odd or disreputable member of a group. This usage derives from the recessive trait that causes an occasional black lamb to be born into an entirely white flock. These black sheep were considered undesirable by shepherds, as black wool is not as commercially viable as white wool. Citizens who accept overbearing governments have been referred to by the Portmanteau word, Portmanteau neologism of sheeple. Somewhat differently, the adjective "sheepish" is also used to describe embarrassment.
In heraldry
In British heraldry, sheep appear in the form of rams, sheep proper and lambs. These are distinguished by the ram being depicted with horns and a tail, the sheep with neither and the lamb with its tail only. A further variant of the lamb, termed the Pascal Lamb (heraldry), Paschal lamb, is depicted as carrying a Christian cross and with a Halo (religious iconography), halo over its head. Rams' heads, portrayed without a neck and facing the viewer, are also found in British armories. The fleece, depicted as an entire sheepskin carried by a ring around its midsection, originally became known through its use in the arms of the Order of the Golden Fleece and was later adopted by towns and individuals with connections to the wool industry.[Arthur Fox-Davies, ''A Complete Guide to Heraldry'', T.C. and E.C. Jack, London, 1909, 211-213, https://archive.org/details/completeguidetoh00foxduoft.] A sheep on a blue field is depicted on the Coat of arms of Denmark, greater/royal arms of the king of Denmark to represent the Faroe Islands. In 2004 a modernized arms has been adopted by the Faroe Islands, which based on a 15th century coat of arms.
Religion and folklore
In antiquity, symbolism involving sheep cropped up in religions in the ancient Near East, the Mideast, and the Mediterranean area: Çatalhöyük, ancient Egyptian religion, the Cana'anite and Phoenician tradition, Judaism, Greek mythology, Greek religion, and others. Religious symbolism and ritual involving sheep began with some of the first known faiths: Skulls of rams (along with bulls) occupied central placement in shrines at the Çatalhöyük settlement in 8,000 BCE. In Ancient Egyptian religion, the ram was the symbol of several gods: Khnum, Heryshaf and Amun (in his incarnation as a god of fertility). Other deities occasionally shown with ram features include the goddess Ishtar, the Phoenician god Baal-Hamon, and the Babylonian god Ea-Oannes. In Madagascar, sheep were not eaten as they were believed to be incarnations of the souls of ancestors.
There are many ancient Greek references to sheep: that of Chrysomallos, the golden-fleeced ram, continuing to be told through into the modern era. Astrology, Astrologically, Aries (astrology), Aries, the ram, is the first sign of the classical Greek zodiac, and the sheep is the eighth of the twelve animals associated with the 12-year cycle of in the Chinese zodiac, related to the Chinese calendar.[ In Mongolia, shagai are an ancient form of dice made from the cuboid bones of sheep that are often used for fortunetelling purposes.
Sheep play an important role in all the Abrahamic faiths; Abraham, Isaac, Jacob, Moses, King David and the Islamic prophet Muhammad were all shepherds. According to the Biblical story of the Binding of Isaac, a ram is sacrificed as a substitute for Isaac after an angel stays Abraham's hand (in the Islamic tradition, Abraham was about to sacrifice Ishmael). Eid al-Adha is a major annual festival in Islam in which sheep (or other animals) are sacrificed in remembrance of this act.] Sheep are occasionally sacrificed to commemorate important secular events in Islamic cultures. Greeks and Romans sacrificed sheep regularly in religious practice, and Judaism once sacrificed sheep as a Korban (sacrifice), such as the Korban Pesach, Passover lamb.[ Ovine symbols—such as the ceremonial blowing of a shofar—still find a presence in modern Judaic traditions.
Collectively, followers of Christianity are often referred to as a flock, with Christ as the The Good Shepherd (Christianity), Good Shepherd, and sheep are an element in the Christian iconography of the Nativity of Jesus in art, birth of Jesus. Some Christian saints are considered Patron saints of occupations and activities, patrons of shepherds, and even of sheep themselves. Christ is also portrayed as the Sacrificial lamb of God (''Agnus Dei'') and Easter celebrations in Greece and Romania traditionally feature a meal of Paschal lamb. A church leader is often called the pastor, which is derived from the Latin word for shepherd. In many western Christian traditions bishops carry a staff, which also serves as a symbol of the episcopal office, known as a crosier, which is modeled on the shepherd's crook.
Sheep are key symbols in fables and nursery rhymes like ''The Wolf in Sheep's Clothing'', ''Little Bo Peep'', ''Baa, Baa, Black Sheep'', and ''Mary Had a Little Lamb''; novels such as George Orwell's ''Animal Farm'' and Haruki Murakami's ''A Wild Sheep Chase''; songs such as Johann Sebastian Bach, Bach's ''Sheep may safely graze'' (''Schafe können sicher weiden'') and Pink Floyd's "Sheep (Pink Floyd song), Sheep", and poems like William Blake's "The Lamb (poem), The Lamb".
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See also
* Chris (sheep)
* Dry Sheep Equivalent
* Fictional sheep
* Sheepfold
* Shrek (sheep)
* Sonny Wool
* U.S. Sheep Experiment Station
* Venray sheep companies
* Sheep–goat hybrid
References
Sources
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External links
American Sheep Industry
(Queensland)
National Sheep Association
(UK)
New Zealand Sheepbreeders Association
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sheep genome
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