''Shamosaurus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of herbivorous basal

ankylosaurid Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...

ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...

from

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

(

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...

stage) deposits of Höövör,

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

Discovery and naming

In 1977, a Soviet-Mongolian expedition discovered the skeleton of an unknown ankylosaurian at the Hamrin-Us site in

Dornogovi Province Dornogovi ( mn, Дорноговь, ''East Gobi'') is one of the 21 aimags (provinces) of Mongolia. It is located in the southeast of the country, bordering PR China's autonomous region of Inner Mongolia. Dornogovi is located in the Gobi desert ...

. This was the first discovery of an ankylosaur in the

Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...

of Mongolia.T.A. Tumanova, 1983, "Pervyy ankilozavr iz nizhnego mela Mongolii", In: L.P. Tatarinov, R. Barsbold, E. Vorobyeva, B. Luvsandanzan, B.A. Trofimov, Yu. A. Reshetov, & M.A. Shishkin (eds.), ''Iskopayemyye reptilii mongolii''. Trudy Sovmestnaya Sovetsko-Mongol'skaya Paleontologicheskaya Ekspeditsiya 24: 110-118 In

1983 The year 1983 saw both the official beginning of the Internet and the first mobile cellular telephone call. Events January * January 1 – The migration of the ARPANET to Internet protocol suite, TCP/IP is officially completed (this is consid ...

Tatyana Tumanova Tatiana (or Tatianna, also romanized as Tatyana, Tatjana, Tatijana, etc.) is a female name of Sabine-Roman origin that became widespread in Eastern Europe. Variations * be, Тацця́на, Tatsiana * bg, Татяна, Tatyana * germa ...

named and described the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

''Shamosaurus scutatus''. The generic name is derived from

Mandarin Mandarin or The Mandarin may refer to: Language * Mandarin Chinese, branch of Chinese originally spoken in northern parts of the country ** Standard Chinese or Modern Standard Mandarin, the official language of China ** Taiwanese Mandarin, Stand ...

''sha mo'', "sand desert", the Chinese name for the

Gobi The Gobi Desert (Chinese: 戈壁 (沙漠), Mongolian: Говь (ᠭᠣᠪᠢ)) () is a large desert or brushland region in East Asia, and is the sixth largest desert in the world. Geography The Gobi measures from southwest to northeast an ...

. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

means "protected by a shield" in

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

, a reference to the body armour. ''Shamosaurus'' is known from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

PIN N 3779/2, collected from the Dzunbain Formation, equivalent to the Khukhtekskaya Formation and dating from the

, about 115 million years old. It consists of a complete

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

, lower

jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...

s and partial postcranial skeleton with armor. Only the skull was described in 1983. Later the specimens PIN 3779/1, a skull piece, and PIN 3101, a lower jaw, were referred. The fossils are part of the collection of the Palaeontological Institute at

Moscow Moscow ( , US chiefly ; rus, links=no, Москва, r=Moskva, p=mɐskˈva, a=Москва.ogg) is the capital and largest city of Russia. The city stands on the Moskva River in Central Russia, with a population estimated at 13.0 million ...

. The holotype skull is exhibited there, together with two cervical halfrings.

Description

''Shamosaurus'' was a medium-sized ankylosaurian. In 2010,

Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...

estimated its body length at 5 metres (16 ft), its weight at 2 tonnes (2.2 short tons). In 2012

Holtz Holtz is the surname of: * Alexander Holtz (born 2002), Swedish ice hockey player * Andrew Holtz, American journalist * Carl Holtz (1920–2006), American oarsman and farmer * Daniel Holtz, a fictional character on the TV series ''Angel'' * Eric H ...

gave a higher estimation of 7 meters (23 ft). In 2014,

Victoria Megan Arbour Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...

gave a revised list of distinguishing traits. The

osteoderms Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct ...

on the skull roof are not very pronounced nor separated as distinctive ''caputegulae'', head tiles. The squamosal horns on the rear skull corners are short and slightly rounded. The quadratojugal horn on the cheek has its apex in the middle. The rear rim of the skull roof has no clear (nuchal) processes.Arbour, Victoria Megan, 2014. ''Systematics, evolution, and biogeography of the ankylosaurid dinosaurs''. Ph.D thesis, University of Alberta ''Shamosaurus scutatus'' shares many

cranial Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

similarities with ''

Gobisaurus ''Gobisaurus'' is an extinct genus of herbivorous basal ankylosaurid ankylosaur from the Upper Cretaceous (and possibly also the Lower Cretaceous) of China (''Nei Mongol Zizhiqu''). The genus is monotypic, containing only the species ''Gobisaur ...

domoculus'', including a rounded

squamosal The squamosal is a skull bone found in most reptiles, amphibians, and birds. In fishes, it is also called the pterotic bone. In most tetrapods, the squamosal and quadratojugal The quadratojugal is a skull bone present in many vertebrates, including ...

, large elliptical

orbital fenestrae The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

(oval eye sockets) and oval external nares (nostrils), a

deltoid Deltoid (delta-shaped) can refer to: * The deltoid muscle, a muscle in the shoulder * Kite (geometry), also known as a deltoid, a type of quadrilateral * A deltoid curve, a three-cusped hypocycloid * A leaf shape * The deltoid tuberosity, a part of ...

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...

profile with a narrow

rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ships * Ros ...

(the snout is tongue-shaped and narrow in top view), quadratojugal protuberances (cheek horns), and caudolaterally directed paroccipital processes (extensions of the rear skull obliquely pointing to behind and sideways). But the two

taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

may be distinguished by differences in the length of the maxillary tooth row (40% instead of 26,7% of total skull length), an unfused basipterygoid-pterygoid process in ''Gobisaurus'', the presence of an elongate vomerine premaxillary process in ''Gobisaurus'', and the presence of cranial sculpting in ''Shamosaurus'', but not in ''Gobisaurus''.Vickaryous M.K., Maryańska T., Weishampel D.B., 2004, "Ankylosauria". Chapter 17 in: Weishampel D.B., Dodson P., Osmólska H., editors. ''The Dinosauria''. 2nd ed. Berkeley (CA): University of California Press. p. 363–392 The last difference was denied by Arbour who concluded that the degree of sculpting was roughly the same. ''Shamosaurus'' had a rather flat skull. The upper beak was sharp and obliquely appending to the front. The beak lacked any teeth. The jaw joint was located far behind the level of the eye socket. The

occipital condyle The occipital condyles are undersurface protuberances of the occipital bone in vertebrates, which function in articulation with the superior facets of the atlas vertebra. The condyles are oval or reniform (kidney-shaped) in shape, and their anteri ...

, and with it the entire head, was obliquely directed to below. A secondary palate was present. The armour of ''Shamosaurus'' contained two cervical halfrings with six segments each, protecting the neck. Also keeled

osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amp ...

s and flat oval scutes were present.

Phylogeny

Tumanova placed ''Shamosaurus'' in the

Ankylosauridae Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...

. She also named a Shamosaurinae, a group of basal ankylosaurids. In 2014, Arbour recovered an evolutionary tree in which ''Shamosaurus'' was the

sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...

of ''Gobisaurus''.

Notes

References

*Matthew K. Vickaryous, Anthony P. Russell, Philip J. Currie, and Xi-Jin Zhao. 2001. A new ankylosaurid (Dinosauria: Ankylosauria) from the Lower Cretaceous of China, with comments on ankylosaurian relationships. Canadian Journal of Earth Sciences/Rev. can. sci. Terre 38(12):1767-1780.

External links