Shadow Bass
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The shadow bass (''Ambloplites ariommus'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
in the sunfish
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
(
Centrarchidae Centrarchidae, better known as sunfishes, is a family of freshwater ray-finned fish belonging to the order Perciformes (formerly belonging to the deprecated order Centrarchiformes), native only to North America. There are eight universally inc ...
) of
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Perciformes. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to southeastern United States of America.


Description

The shadow bass, has a
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
containing 36 to 42 scales. There are 11 spines and 11-12 soft rays in the
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
while the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
contains 6-7 spines and 10-11 soft rays. It has a large mouth with the
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
at the level of the centre of its eye. They are olive to light brown in color and there are frequently 2-4 dark brown vertical bands and blotches along their flanks. The dorsal, anal and
caudal fins Fins are distinctive anatomical features composed of bony spine (zoology), spines or Ray (fish fin anatomy), rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bon ...
are usually transparent to light yellow and are mottled with black and gray. The iris is reddish, and the pupil is deep blue. There is a black band which runs from beneath the eye onto the gill cover. The spots on the scales of the shadow bass are triangular while the similar
rock bass The rock bass (''Ambloplites rupestris''), also known as the rock perch, goggle-eye, red eye, and black perch, is a freshwater fish native to east-central North America. This red eyed creature is a species of freshwater fish in the sunfish fa ...
has rectangular spots on its scales. The shadow bass is a medium sized sunfish which has an average
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
of around and a wmaximum length of and weight of . Females normally live longer than the males but the males are faster growing and grow larger.


Geographic distribution

The shadow bass' native range includes the southeastern United States from the Apalachicola River drainage in Georgia to the lower Mississippi basin in
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
. The two isolated populations in the St. Francis, Black Arkansas Red, and upper Ouachita River drainages in
Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ...
and
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...
in the Ozark regions. In
Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...
, the shadow bass can be found in most drainages south of the
Tennessee River The Tennessee River is the largest tributary of the Ohio River. It is approximately long and is located in the southeastern United States in the Tennessee Valley. The river was once popularly known as the Cherokee River, among other names, ...
. Other river drainages that shadow bass inhabit are Blackwater and
Chattahoochee River The Chattahoochee River forms the southern half of the Alabama and Georgia border, as well as a portion of the Florida - Georgia border. It is a tributary of the Apalachicola River, a relatively short river formed by the confluence of the Chatta ...
s in southern Georgia and northern
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
. The apparent absence of shadow bass in
Black Belt Black Belt may refer to: Martial arts * Black belt (martial arts), an indication of attainment of expertise in martial arts * ''Black Belt'' (magazine), a magazine covering martial arts news, technique, and notable individuals Places * Black B ...
streams is probably the result of low stream flow and poor water-quality conditions during the late summer when temperatures are high. Channelization is a possible factor in the distribution of the shadow bass in
Mississippi River The Mississippi River is the second-longest river and chief river of the second-largest drainage system in North America, second only to the Hudson Bay drainage system. From its traditional source of Lake Itasca in northern Minnesota, it f ...
systems, but no difference in the distribution of shadow bass was found.


Ecology

The shadow bass is a freshwater fish that inhabits a climate range of 38 to 29°N in the United States. It prefers slow-flowing water over substrates of silt-free mud, sand, gravel, and pebbles. Brushy and vegetated pools are preferred microhabitat in most streams. Undercut banks and woody debris in small to medium-sized rivers and streams are also preferred habitat. The quantitative diet of shadow bass consists mostly of small invertebrates, mainly
crayfish Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, craydids, crawdaddies, crawdads, freshwater lobsters, mountain lobsters, rock lobsters, mu ...
when less than three inches; after three inches, they can begin feeding on small fish species such as
darters The darters, anhingas, or snakebirds are mainly tropical waterbirds in the family Anhingidae, which contains a single genus, ''Anhinga''. There are four living species, three of which are very common and widespread while the fourth is rarer and ...
, madtoms, and minnows. The main predator and competitor of the shadow bass is the smallmouth bass which inhabits the same microhabitat. Research on the introduction of
rainbow trout The rainbow trout (''Oncorhynchus mykiss'') is a species of trout native to cold-water tributaries of the Pacific Ocean in Asia and North America. The steelhead (sometimes called "steelhead trout") is an anadromous (sea-run) form of the coasta ...
(''Oncorhynchus mykiss'') in shadow bass habitat to measure competition between the species, the data showed no effect on shadow bass populations. Factors by humans that might attribute to population decline of the shadow bass channelization, sedimentation from logging, and habitat encroachment.


Life history

Shadow bass normally
spawn Spawn or spawning may refer to: * Spawn (biology), the eggs and sperm of aquatic animals Arts, entertainment, and media * Spawn (character), a fictional character in the comic series of the same name and in the associated franchise ** '' Spawn: Ar ...
from January to August, spawning season being determined by water temperature and so by location to somne extent too. Spawning is earlier in southern Alabama or southern Georgia as the wate is warmer in these regions, while fish found at higher latitudes typically spawn later when the water reached the ideal spawning temperature of .The breeding males show obvious darkening to the membranes of the
pelvic The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...
and anal fins while these remain yellow or white in the females. Sexual maturity is attained at a year old and a mature female may lay 1,33 eggs in a spawning season, although after cold winters fewer, larger eggs are laid and this means that spawn hatching rates are not affected by temperature. This strategy of laying a smaller number of larger eggs after colder winters or a larger number of smaller eggs after milder winters ensures that the population of shadow bass has a consistent recruitment rate.


Taxonomy and etymology

The shadow bass is a member of the centrarchids or sunfish family and is closely related to the rock bass ''Ambloplites rupestris'' along with one more related species occurring naturally. The shadow bass was previously considered to be a subspecies of the rock bass (''A. rupestris''). It was first formally described in 1936 by Peter Viosca Jr. with the type locality given as Little Bogue Falaya Creek, 3 miles north of
Covington, Louisiana Covington is a city in, and the parish seat of, St. Tammany Parish, Louisiana, United States. The population was 11,564 at the 2020 United States census. It is located at a fork of the Bogue Falaya and the Tchefuncte River. Covington is part of ...
. The
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name is derived from Greek and is a compound of ''ambloo'' meaning "blunt" and ''oplites'' meaning "shield", while the
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
means "large-eyed".


Relationship with humans


Management

Through its range even for the two isolated populations, the shadow bass is not listed as being in any danger, and the shadow bass is federally listed as G5 secure, while the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
classify its status as
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
. Current management for the shadow bass in the rivers in Mississippi was to protects pools and prevent sediment pollution in areas of shadow bass populations. In most places, shadow bass are not considered a game fish and receive little management attention. In Missouri, however, where shadow bass has a demand in the game fish market, biologists are conducting population studies using mark and recapture methods and information for anglers to provide data to determine what, if any, angling restrictions need to apply to shadow bass fishing.


Angling

The IGFA all tackle world record for the shadow bass stands at 0.82 kg (1 lb 13oz) caught from the Spring River in
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...
.


References


Bibliography

*
FishBase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.

''Ambloplites ariommus''
* {{Taxonbar, from=Q3768484 Ambloplites Freshwater fish of the United States Fish described in 1936