Setophaga Ruticilla
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The American redstart (''Setophaga ruticilla'') is a New World warbler. It is unrelated to the Old World (common) redstart.


Taxonomy

The American redstart was described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in 1758 in the
10th edition 1 (one, unit, unity) is a number representing a single or the only entity. 1 is also a numerical digit and represents a single unit of counting or measurement. For example, a line segment of ''unit length'' is a line segment of length 1. I ...
of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Motacilla ruticilla''. The genus name ''Setophaga'' is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
σής : ''sēs'' (genitive σητός : sētós) "moth", and ', "eating". The specific ''ruticilla'' is New Latin for "redstart" from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''rutilus'', "red", and New Latin ''cilla'', "tail". "Redstart" refers to the male's red tail, "start" being an old word for tail.


Description

The American redstart is a smallish warbler. It measures in total length and has a wingspan of . Its length is boosted by a relatively long tail and it is one of the lightest birds in its family. Weight is considerably less in winter than in summer. Males weigh an average of in summer but drop to in winter, while females drop even more from an average of to an average of . Among standard measurements, the wing chord is , the
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...
is , the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
is and the tarsus is . The breeding males are unmistakable, jet black above apart from large orange-red patches on their wings and tails. Their breast sides are also orange, with the rest of their underparts white. In their other plumages, American redstarts display green in their upperparts, along with black central tails and grey heads. The orange patches of the breeding males are replaced by yellow in the plumages of the females and young birds. Orange and yellow coloration is due to the presence of
carotenoid Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic compound, organic pigments that are produced by plants and algae, as well as several bacteria, and Fungus, fungi. Carotenoids give the characteristic color to pumpki ...
s; males possess the red carotenoid canthaxanthin and the yellow carotenoids canary xanthophyll A and B, all of which mix together to produce an orange color, while the females possess only the yellow carotenoids. Recent research indicates that an age and sex effect on observed color attributes of hue, brightness, and saturation exists in American redstarts, with the exception for saturation, which only showed an age effect. Their song is a series of musical ''see'' notes. Their call is a soft ''chip''.


Distribution and habitat

Although perhaps not as common as in the past, the redstart appears to be one of the most stable and abundant species of New World warbler; its numbers exceeded in total by the
common yellowthroat The common yellowthroat (''Geothlypis trichas'') is a New World warbler. In the U.S. Midwest, it is also known as the yellow bandit. It is an abundant breeder in North America, ranging from southern Canada to central Mexico. The genus name ''Geot ...
,
yellow warbler The yellow warbler (''Setophaga petechia'') is a New World warbler species. Yellow warblers are the most widespread species in the diverse genus ''Setophaga'', breeding in almost the whole of North America, the Caribbean, and down to northern S ...
and
yellow-rumped warbler The yellow-rumped warbler (''Setophaga coronata'') is a regular North American bird species that can be commonly observed all across the continent. Its extensive distribution range connects both the Pacific and Atlantic coasts of the U.S. as well ...
, because of much wider natural breeding ranges in those species and perhaps exceeding those in sheer density within appropriate range. Their breeding is in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, spanning southern
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
and the eastern United States. They are migratory, wintering in
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, the
West Indies The West Indies is a subregion of North America, surrounded by the North Atlantic Ocean and the Caribbean Sea that includes 13 independent island countries and 18 dependencies and other territories in three major archipelagos: the Greater A ...
, and northern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, and are very rare vagrants to western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. During the breeding season, the redstart inhabits open-canopy, mostly deciduous forests, second growth, and forest edges. It is insectivorous, often sharing its foraging habitats with other warblers, and is found feeding in the mid to lower regions of a tree or shrub. A wide range of habitats are occupied during migration, including many shrubby areas. On its wintering grounds in Central and South America, the redstart may be found in nearly all woody habitats but tends to avoid non-forested agricultural areas. It is often found in
shade-grown coffee Shade-grown coffee is a form of the crop produced from coffee plants grown under a canopy of trees. A canopy of assorted types of shade trees is created to cultivate shade-grown coffee. Because it incorporates principles of natural ecology to pro ...
plantations, which provide native trees and shrubs, as well as coffee trees. Elevations occupied vary by location, with redstarts found at elevations up to in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, but only in
Jamaica Jamaica (; ) is an island country situated in the Caribbean Sea. Spanning in area, it is the third-largest island of the Greater Antilles and the Caribbean (after Cuba and Hispaniola). Jamaica lies about south of Cuba, and west of His ...
.


Behavior


Breeding

The breeding habitats of the redstarts are open woodlands or scrub, often located near water. They nest in the lower part of a bush, laying 2–5 eggs in a neat cup-shaped nest. The clutch is incubated by the female for 10 to 13 days. The young fledge after 9 days in the nest, and may remain with one parent for up to 3 weeks afterwards. First-year males are able to reproduce during their first breeding season, but they retain the female-like plumage which may contribute to low reproductive success (less than 50% of first-year males) until year 2. In contrast, most first-year females successfully reproduce during their first breeding season. There is evidence for a skewed sex ratio that results in a surplus of unmated males.Sherry, T., R. Holmes. 1997. ''American Redstart (Setophaga ruticilla)'' (On-line). The Birds of North America Online. American redstarts display a mixed mating strategy; they are predominantly
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
but around 25% of males maintain multiple territories and are
polygynous Polygyny (; from Neoclassical Greek πολυγυνία (); ) is the most common and accepted form of polygamy around the world, entailing the marriage of a man with several women. Incidence Polygyny is more widespread in Africa than in any ...
. Even within monogamous pairs, a high proportion of offspring—as many as 40%—are not fathered by the male of the pair. The intensity of the male's coloration (which is due to
carotenoid Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic compound, organic pigments that are produced by plants and algae, as well as several bacteria, and Fungus, fungi. Carotenoids give the characteristic color to pumpki ...
pigments) predicts their success at holding territory in their non-breeding, winter locations in the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
, the probability that they will be polygynous, and the proportion of offspring in their nests that they will themselves father. Males are invariably very territorial and the superior males occupy the best habitats, such as moist mangroves, while inferior males occupy secondary habitats such as dry scrub forests.


Feeding

The redstarts feed almost exclusively on
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s which are usually caught by flycatching. American redstarts also have been known to catch their insect prey by gleaning it from leaves. This is a very active species. The tail is often held partly fanned out. These birds have been observed flashing the orange and yellow of their tails on and off to startle and chase insects from the underbrush. Overall, this species is a very flexible, opportunistic feeder that can easily adapt to varying habitat, season, insect community, vegetation structure, and time of day. The diet consists largely of
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s,
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s, flies,
leafhopper A leafhopper is the common name for any species from the family Cicadellidae. These minute insects, colloquially known as hoppers, are plant feeders that suck plant sap from grass, shrubs, or trees. Their hind legs are modified for jumping, and a ...
s and
planthopper A planthopper is any insect in the infraorder Fulgoromorpha, in the suborder Auchenorrhyncha, a group exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment ...
s, small
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
s,
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s, aphids,
stoneflies Plecoptera is an order of insects, commonly known as stoneflies. Some 3,500 species are described worldwide, with new species still being discovered. Stoneflies are found worldwide, except Antarctica. Stoneflies are believed to be one of the mo ...
and
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s. Few berries and seeds are consumed, but are most often from
barberry ''Berberis'' (), commonly known as barberry, is a large genus of deciduous and evergreen shrubs from tall, found throughout temperate and subtropical regions of the world (apart from Australia). Species diversity is greatest in South America a ...
,
serviceberry ''Amelanchier'' ( ), also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry (or just sarvis), juneberry, saskatoon, sugarplum, wild-plum or chuckley pear,A Digital Flora of Newfoundland and Labrador Vascular Plants/ref> is a g ...
, and
magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendro ...
.


Mortality

The oldest known banded redstart lived to over 10 years of age. Other adults have been known to reach around 5 years. However, few survive past the first stages of life, as the bird is vulnerable to both terrestrial and aerial predators. Highest rates of predation occur during the breeding season when eggs and helpless nestlings are abundant and easy prey for varied predators. Females mostly brood during this period and thus often fall prey to nest predators. Common terrestrial predators include
red squirrel The red squirrel (''Sciurus vulgaris'') is a species of tree squirrel in the genus ''Sciurus'' common throughout Europe and Asia. The red squirrel is an arboreal, primarily herbivorous rodent. In Great Britain, Ireland, and in Italy numbers ...
s,
fisher Fisher is an archaic term for a fisherman, revived as gender-neutral. Fisher, Fishers or The Fisher may also refer to: Places Australia *Division of Fisher, an electoral district in the Australian House of Representatives, in Queensland *Elect ...
s, eastern chipmunks,
American black bear The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
s,
flying squirrel Flying squirrels (scientifically known as Pteromyini or Petauristini) are a tribe of 50 species of squirrels in the family Sciuridae. Despite their name, they are not in fact capable of full flight in the same way as birds or bats, but they ar ...
s, fox snakes, and
domestic cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members o ...
s. Aerial predators take nestlings, eggs, or even adults in flight. Aerial predators include jaegers,
blue jay The blue jay (''Cyanocitta cristata'') is a passerine bird in the family Corvidae, native to eastern North America. It lives in most of the eastern and central United States; some eastern populations may be migratory. Resident populations are a ...
s,
common raven The common raven (''Corvus corax'') is a large all-black passerine bird. It is the most widely distributed of all corvids, found across the Northern Hemisphere. It is a raven known by many names at the subspecies level; there are at least e ...
s, northern saw-whet owls, common grackles,
northern goshawk The northern goshawk (; ''Accipiter gentilis'') is a species of medium-large bird of prey, raptor in the Family (biology), family Accipitridae, a family which also includes other extant diurnal raptors, such as eagles, buzzards and harrier (bird) ...
s, sharp-shinned hawks, and Cooper's hawks.


Conservation

Successful conservation efforts of the redstart, as for any other migrating bird, include protecting and providing habitat throughout its entire range. The benefits to coffee farms that redstarts and other "coffee birds" provide have encouraged coffee farmers to adapt shade trees and adjacent forest patches in their farming practices as additional habitat for the birds. While shade tree coffee farms offer a somewhat practical compromise between habitat preservation and agriculture, there is still not enough data to back the proposition that practices like shade tree coffee farms can replace the natural habitat that was once there. Still, the most effective method for American redstart conservation would be natural habitat preservation at wintering and breeding grounds.


References


External links


American redstart species account
– Cornell Lab of Ornithology

- USGS Patuxent Bird Identification InfoCenter
American redstart bird sound
at Florida Museum of Natural History * {{Taxonbar, from=Q922772 American redstart Birds of North America Birds of the Dominican Republic American redstart American redstart