Semnornis
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The toucan-barbets are the small
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Semnornis''. This was often included in the
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
barbets but recently usually considered a distinct
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Semnornithidae; alternatively, all barbets might be moved to the
toucan Toucans (, ) are members of the Neotropical near passerine bird family Ramphastidae. The Ramphastidae are most closely related to the American barbets. They are brightly marked and have large, often colorful bills. The family includes five ge ...
family
Ramphastidae Toucans (, ) are members of the Neotropical near passerine bird family Ramphastidae. The Ramphastidae are most closely related to the American barbets. They are brightly marked and have large, often colorful bills. The family includes five gen ...
as a subfamily, Semnornithinae. It contains only two
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, the
toucan barbet The toucan barbet (''Semnornis ramphastinus'') is a barbet native to western Ecuador and Colombia. Along with the prong-billed barbet, it forms the family Semnornithidae, and is closely related to the toucans. It is a medium-sized barbet with a ...
(''S. ramphastinus'') and the
prong-billed barbet The prong-billed barbet (''Semnornis frantzii'') is a distinctive, relatively large-billed bird native to humid highland forest of Costa Rica and western Panama. Taxonomy and systematics The prong-billed barbet was traditionally placed in fam ...
(''S. frantzii'').


Description

The ''Semnornis'' barbets are fairly large barbets, measuring between . The toucan barbet is larger than the prong-billed barbet and considerably heavier. They possess large, swollen bills and lack strong
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
in their
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
. The plumage of the prong-billed barbet is orange-brown, and that of the toucan barbet is more distinctively patterned with black, red, grey and gold.


Distribution and habitat

The ''Semnornis'' toucan-barbets are found in the
Neotropic The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
s. The prong-billed barbet is restricted to the humid highland forests of
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
and
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
. The toucan barbet is found in similar habitats in the western montane forests of
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
and
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
. In addition to primary forest they may occupy forest edges and secondary growth. Neither species is migratory, and young birds do not appear to disperse very far after fledging; young toucan barbets only disperse 0.5 km.


Behaviour

The Semnornithidae are highly social, and may be seen either in small groups of up to five or six individuals, or as singles. They are active during the day and are early risers. The prong-billed barbet sleeps in communal roosts at night in the non-breeding season. As many as 19 birds may roost together in a hole, either a modified nest or the abandoned nest of a
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. ...
. During the breeding season pairs roost in their own nests.


Diet and feeding

The diet of these two species is made up of fruits and insects. The ratio of the two is more similar to the toucans than other barbets and is dominated by fruits. A 1993 study of the stomach contents of these two species found only fruit. Fruits may be eaten whole, held in the foot and broken and eaten, or crushed and only the juices eaten. Insects are more common in the diet of nestlings, and compose 40% of the food brought to the nest in toucan-barbets. Toucan-barbets may also feed their chicks small numbers of vertebrates. They have also been recorded eating flowers.


Breeding

Both species of toucan-barbet are
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
breeders. Prong-billed barbets defend breeding
territories A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
from all others of their species. Toucan barbets, on the other hand, have territories but are helped in raising the young by helpers.


Species


References


Further reading

*''Edinburgh New Philosophical Journal, (New series)'' 2 no.2 p. 404


External links


Barbet videos
on the Internet Bird Collection

{{Taxonbar, from=Q204765   Bird genera Higher-level bird taxa restricted to the Neotropics Taxa named by Charles Wallace Richmond