Semiothisa Clathrata
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The latticed heath (''Chiasmia clathrata'') is a
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
of the family Geometridae, belonging to the subfamily Ennominae, placed in the tribe
Macariini The Macariini are a tribe of geometer moths in the subfamily Ennominae. Though they share many traits with the Sterrhinae, this is probably plesiomorphic rather than indicative of a close relationship, and DNA sequence data points to the Boarmi ...
. The genus was erected by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his 1758 10th edition of ''Systema Naturae''.


Taxonomy

The current placement of the latticed heath in the genus ''
Chiasmia ''Chiasmia'' is a genus of moths in the family Geometridae. It was described by Jacob Hübner Jacob Hübner (20 June 1761 – 13 September 1826, in Augsburg) was a German entomologist. He was the author of ''Sammlung Europäischer Schmetterling ...
'' follows from the revision by Malcolm J. Scoble (2002) of the tribe Macariini, when he showed that true ''
Semiothisa ''Semiothisa'' is a genus of moths in the family Geometridae. It was erected by Jacob Hübner in 1818. Species Some species of this genus are: * '' Semiothisa albivia'' (Prout, 1915) * '' Semiothisa arenosa'' (Butler, 1875) * '' Semiothisa assi ...
'' species were restricted to the Americas. There are a number of described subspecies. Molecular work has confirmed the placement of the species within ''Chiasmia''.


Subspecies

Subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
include: * ''Chiasmia clathrata clathrata'' (Linné) * ''Chiasmia clathrata centralasiae'' (Krulikowski, 1911) * ''Chiasmia clathrata djakonovi'' (Kardakoff, 1928) * ''Chiasmia clathrata kurilata'' (Bryk, 1942)


Distribution and habitat

This species can be found throughout Europe, the
Near East The ''Near East''; he, המזרח הקרוב; arc, ܕܢܚܐ ܩܪܒ; fa, خاور نزدیک, Xāvar-e nazdik; tr, Yakın Doğu is a geographical term which roughly encompasses a transcontinental region in Western Asia, that was once the hist ...
,
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
, and east through
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
,
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
,
Amur River The Amur (russian: река́ Аму́р, ), or Heilong Jiang (, "Black Dragon River", ), is the world's List of longest rivers, tenth longest river, forming the border between the Russian Far East and Northeast China, Northeastern China (Inne ...
, northern
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
,
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. It is a fairly common species in the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
.Skinner, B., 1984. ''Colour Identification Guide to Moths of the British Isles'' These moths inhabit a range of open areas, including grassland, moorland, and waste ground.


Description


Adult

''Chiasmia clathrata'' has a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
of 20–25 mm.''UKMoths''
/ref> In this rather variable species the background colour of the wings varies from yellowish to white, with a network of brown lines (hence its common name). These lines vary in thickness and sometimes the wings are almost entirely dark brown. File:Chiasmia clathrata - Géomètre à barreaux MHNT.jpg , Female – dorsal and ventral sides File: Chiasma clathrata02.jpg, Spread wings File: Semiothisa clathrata 02.jpg, Underside, at night


Larva

The final
instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or ass ...
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
is pale green with white lines, including a strong lateral line and thin dorsal lines along the body. Abdominal segments A1 through A5 have a strong white line across the rear end of each segment that ends just above the lateral line.Porter, J., 1997. ''The Colour Identification Guide to Caterpillars of the British Isles''. Viking Press, Harmondsworth, Middlesex. xii + 275 pp.


Ecology

In the British Isles there are one or two generations annually, with adults seen at any time from May to September. These moths are diurnal but they can also be observed at night where they are attracted to artificial light.Truxa, C., Fiedler, K. (2012), Attraction to light - from how far do moths (Lepidoptera) return to weak artificial sources of light? ''European Journal of Entomology'', 109 (1):77-84. Larvae feed on bedstraws (''
Galium mollugo ''Galium mollugo'', common name hedge bedstraw or false baby's breath, is a herbaceous perennial plant of the family Rubiaceae. It shares the name hedge bedstraw with the related European species, Galium album. Description ''Galium mollugo'' can ...
'', ''
Galium verum ''Galium verum'' (lady's bedstraw or yellow bedstraw) is a herbaceous perennial plant of the family Rubiaceae. It is widespread across most of Europe, North Africa, and temperate Asia from Israel, Lebanon and Turkey to Japan and Kamchatka. It is ...
'') and various
legumes A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock fo ...
such as clovers (''
Trifolium medium ''Trifolium medium'', the zigzag clover, is a flowering plant species in the bean family Fabaceae. It is similar in appearance to red clover, ''Trifolium pratense'', but the leaflets are narrower and have no white markings and the narrow stipules ...
'', '' Trifolium pratense''), trefoils, lucerne ('' Medicago sativa'') and
meadow vetchling ''Lathyrus pratensis'' or meadow vetchling, yellow pea, meadow pea and meadow pea-vine, is a perennial legume that grows to 1.2 m in height. The hermaphrodite flowers are pollinated by bees. As a perennial, this plant reproduces itself over man ...
, primarily in June and July and from mid-August through September, though in Ireland and northern Britain larvae occur in July and August. The species overwinters as a
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
.


Bibliography

* , 1986. ''Collins Guide to the Insects of Britain and Western Europe'' (Reprinted 1991) * , 2011. Phylogenetic relationships of selected European Ennominae (Lepidoptera: Geometridae). ''European Journal of Entomology'' 108: 267–273. * , 1997. ''The Colour Identification Guide to Caterpillars of the British Isles''. Viking Press, Harmondsworth, Middlesex. xii + 275 pp. * , 1984. ''Colour Identification Guide to Moths of the British Isles'' * , 2002. A review of the genera of Macariini with a revised classification of the tribe (Geometridae: Ennominae). ''Zoological Journal of the Linnean Society'' 134 (3): 257–315.


References


External links

* Paolo Mazzei, Daniel Morel, Raniero Panfil
''Moths and Butterflies of Europe and North Africa''''Lepiforum e.V.''
*Heiner Ziegle
''Schmetterlinge der Schweiz''
{{Taxonbar, from=Q25779 Macariini Moths described in 1758 Moths of Asia Moths of Europe Moths of Africa Taxa named by Carl Linnaeus Articles containing video clips