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''Chondrus crispus''—commonly called Irish moss or carrageen moss (
Irish Irish may refer to: Common meanings * Someone or something of, from, or related to: ** Ireland, an island situated off the north-western coast of continental Europe ***Éire, Irish language name for the isle ** Northern Ireland, a constituent unit ...
''carraigín'', "little rock")—is a species of
red algae Red algae, or Rhodophyta (, ; ), are one of the oldest groups of eukaryotic algae. The Rhodophyta also comprises one of the largest phyla of algae, containing over 7,000 currently recognized species with taxonomic revisions ongoing. The majority ...
which grows abundantly along the rocky parts of the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
coast of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. In its fresh condition this
protist A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exc ...
is soft and cartilaginous, varying in color from a greenish-yellow, through red, to a dark purple or purplish-brown. The principal constituent is a mucilaginous body, made of the polysaccharide
carrageenan Carrageenans or carrageenins ( ; ) are a family of natural linear sulfated polysaccharides that are extracted from red edible seaweeds. Carrageenans are widely used in the food industry, for their gelling, thickening, and stabilizing properties. T ...
, which constitutes 55% of its dry weight. The organism also consists of nearly 10% dry weight protein and about 15% dry weight
mineral In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid chemical compound with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. ( ...
matter, and is rich in
iodine Iodine is a chemical element with the symbol I and atomic number 53. The heaviest of the stable halogens, it exists as a semi-lustrous, non-metallic solid at standard conditions that melts to form a deep violet liquid at , and boils to a vi ...
and
sulfur Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...
. When softened in
water Water (chemical formula ) is an inorganic, transparent, tasteless, odorless, and nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living organisms (in which it acts as a ...
it has a
sea The sea, connected as the world ocean or simply the ocean, is the body of salty water that covers approximately 71% of the Earth's surface. The word sea is also used to denote second-order sections of the sea, such as the Mediterranean Sea, ...
-like odour and because of the abundant cell wall polysaccharides it will form a jelly when boiled, containing from 20 to 100 times its weight of
water Water (chemical formula ) is an inorganic, transparent, tasteless, odorless, and nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living organisms (in which it acts as a ...
.


Description

''Chondrus crispus'' is a relatively small sea alga, reaching up to a little more than 20 cm in length. It grows from a discoid holdfast and branches four or five times in a dichotomous, fan-like manner. The morphology is highly variable, especially the broadness of the thalli. The branches are 2–15 mm broad and firm in texture, and the color ranges from bright green towards the surface of the water, to deep red at greater depths. The
gametophyte A gametophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has on ...
s (see below) often show a blue
iridescence Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear to gradually change color as the angle of view or the angle of illumination changes. Examples of iridescence include soap bubbles, feathers, butterfl ...
at the tip of the fronds and fertile sporophytes show a spotty pattern. ''
Mastocarpus stellatus ''Mastocarpus stellatus,'' commonly known as carrageenan moss or false Irish moss, is a species in the Rhodophyceae division, a red algae seaweed division, and the Phyllophoracea family. ''M. stellatus'' is closely related to Irish Moss (''Cho ...
'' (Stackhouse) Guiry is a similar species which can be readily distinguished by its strongly channelled and often somewhat twisted thalli.


Distribution

''Chondrus crispus'' is common all around the shores of
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
and can also be found along the coast of Europe including
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its s ...
, the
Faroe Islands The Faroe Islands ( ), or simply the Faroes ( fo, Føroyar ; da, Færøerne ), are a North Atlantic island group and an autonomous territory of the Kingdom of Denmark. They are located north-northwest of Scotland, and about halfway bet ...
. western
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and from ...
to southern
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
. It is found on the Atlantic coasts of
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
and recorded from
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. However, any distribution outside the Northern Atlantic needs to be verified. There are also other species of the same genus in the Pacific Ocean, for example, ''C. ocellatus'' Holmes, ''C. nipponicus'' Yendo, ''C. yendoi'' Yamada ''et'' Mikami, ''C. pinnulatus'' (Harvey) Okamura and ''C. armatus'' (Harvey) Yamada ''et'' Mikami.Hu, Z., Critchley, A.T., Gao T, Zeng X, Morrell, S.L. and Delin, D. 2007 Delineation of Chondrus (Gigartinales, Florideophyceae) in China and the origin of C. crisps inferred from molecular data. Marine Biology Research, 3: 145-154


Ecology

''Chondrus crispus'' is found growing on rock from the middle intertidal
zone Zone or The Zone may refer to: Places Climate and altitude zones * Death zone (originally the lethal zone), altitudes above a certain point where the amount of oxygen is insufficient to sustain human life for an extended time span * Frigid zone, ...
into the subtidal zone, all the way to the ocean floor. It is able to survive with minimal sunlight. ''C. crispus'' is susceptible to infection from the
oomycete Oomycota forms a distinct phylogenetic lineage of fungus-like eukaryotic microorganisms, called oomycetes (). They are filamentous and heterotrophic, and can reproduce both sexually and asexually. Sexual reproduction of an oospore is the resul ...
''
Pythium porphyrae ''Pythium porphyrae'', is a parasitism, parasitic species of oomycete in the family Pythiaceae. It is the cause of red rot disease or red wasting disease, also called ' () in Japanese language, Japanese. The Botanical name, specific epithet '' ...
''.


Uses

''C. crispus'' is an industrial source of
carrageenan Carrageenans or carrageenins ( ; ) are a family of natural linear sulfated polysaccharides that are extracted from red edible seaweeds. Carrageenans are widely used in the food industry, for their gelling, thickening, and stabilizing properties. T ...
commonly used as a thickener and stabilizer in milk products, such as
ice cream Ice cream is a sweetened frozen food typically eaten as a snack or dessert. It may be made from milk or cream and is flavoured with a sweetener, either sugar or an alternative, and a spice, such as cocoa or vanilla, or with fruit such as str ...
and processed foods. In Europe, it is indicated as E407 or E407a. It may also be used as a thickener in calico printing and paper marbling, and for fining
beer Beer is one of the oldest and the most widely consumed type of alcoholic drink in the world, and the third most popular drink overall after water and tea. It is produced by the brewing and fermentation of starches, mainly derived from ce ...
. Irish moss is frequently used with ''Mastocarpus stellatus'' (''Gigartina mamillosa''), ''Chondracanthus acicularis '' (''G. acicularis''), and other
seaweed Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of '' Rhodophyta'' (red), ''Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ...
s, which are all commonly found growing together. Carrageenan may be extracted from tropical seaweeds of the genera ''
Kappaphycus ''Kappaphycus'' is a genus of red algae. Species are distributed in the waters of East Africa, Indonesia, Malaysia, Hainan Island, the Philippines, and Micronesia.Guiry, M. D. In: Guiry, M. D. & G. M. Guiry. 2013''Kappaphycus''.AlgaeBase. Nationa ...
'' and ''
Eucheuma ''Eucheuma'', commonly known as gusô (), is a rhodophyte seaweed that may vary in color (red, brown, and green). ''Eucheuma'' species are used in the production of carrageenan, an ingredient for cosmetics, food processing, and industrial manufac ...
''.


Life history

Irish moss undergoes an alternation of generation lifecycle common in many species of algae (see figure below). The two distinct stages are the sexual
haploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
gametophyte A gametophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has on ...
stage and the asexual
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
sporophyte A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote pr ...
stage. In addition, a third stage - the
carposporophyte {{Short pages monitor