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''Scolopendra heros'', commonly known as the giant desert centipede, giant Sonoran centipede, Texas redheaded centipede, and giant redheaded centipede, is a species of North American centipede found in the Southwestern United States and
Northern Mexico Northern Mexico ( es, el Norte de México ), commonly referred as , is an informal term for the northern cultural and geographical area in Mexico. Depending on the source, it contains some or all of the states of Baja California, Baja California ...
.


Description

''S. heros'' is the largest centipede in North America. It has an average length of , but can reach up to in the wild, and even longer in captivity. Its trunk bears either 21 or 23 pairs of legs. It is aposematically colored, to warn off potential
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s, and a number of color variants are known in the species. The ''castaneiceps'' variant found in Arkansas, Missouri, Texas, and other nearby areas is commonly known as the "giant redheaded centipede" or "Texas redheaded centipede" because of its distinct red head and greenish black body and tail.


Distribution and ecology

''S. heros'' is found in northern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
and the southwestern
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
, from
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ke ...
and
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
in the west to
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the O ...
,
Missouri Missouri is a state in the Midwestern region of the United States. Ranking 21st in land area, it is bordered by eight states (tied for the most with Tennessee): Iowa to the north, Illinois, Kentucky and Tennessee to the east, Arkansas t ...
, and
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
in the east. Although this species is commonly referred to as the "giant desert centipede" because of its presence in the Sonoran and
Chihuahuan Desert The Chihuahuan Desert ( es, Desierto de Chihuahua, ) is a desert ecoregion designation covering parts of northern Mexico and the southwestern United States. It occupies much of far West Texas, the middle to lower Rio Grande Valley and the lo ...
s, and other
arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ...
and
semi-arid A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi- ...
habitats, ''S. heros'' is also found in rocky woodland areas, such as in Arkansas. It remains underground on warm days, emerging in cloudy weather.


Hunting and diet

''S. heros'' is primarily a nocturnal predator and hunts invertebrates and small vertebrates, including rodents, reptiles, and amphibians. It is capable of reaching into the air to grab small flying insects. The centipede uses its venom to subdue prey.


Life cycle

''S. heros'' hatch from eggs. As they grow and mature, like all arthropods they shed and molt away their exoskeleton. Each time they molt they enter a new stage of its life cycle called an instar. Like all Scolopendromorph centipedes, the number of segments they possess remains the same throughout their life. ''S. heros'' is a slow-growing species capable of living over a decade.


Venom

The venom of ''S. heros'' is similar in composition to the venom of other ''Scolopendra'' species, including components such as serotonin,
histamine Histamine is an organic nitrogenous compound involved in local immune responses, as well as regulating physiological functions in the gut and acting as a neurotransmitter for the brain, spinal cord, and uterus. Since histamine was discovered ...
,
lipid Lipids are a broad group of naturally-occurring molecules which includes fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids includ ...
s,
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
s (including
cardiotoxic Cardiotoxicity is the occurrence of heart dysfunction as electric or muscle damage, resulting in heart toxicity. The heart becomes weaker and is not as efficient in pumping blood. Cardiotoxicity may be caused by chemotherapy (a usual example is th ...
proteins and
enzyme Enzymes () are proteins that act as biological catalysts by accelerating chemical reactions. The molecules upon which enzymes may act are called substrates, and the enzyme converts the substrates into different molecules known as products ...
s such as
hemolytic Hemolysis or haemolysis (), also known by several other names, is the rupturing (lysis) of red blood cells (erythrocytes) and the release of their contents (cytoplasm) into surrounding fluid (e.g. blood plasma). Hemolysis may occur in vivo o ...
phospholipase A), and other substances. The mixture is known to act as a
cytolysin Cytolysin refers to the substance secreted by microorganisms, plants or animals that is specifically toxic to individual cells, in many cases causing their dissolution through lysis. Cytolysins that have a specific action for certain cells are n ...
, compromising cellular membranes and rupturing cells. ''S. heros'' venom also contains toxins targeted to its prey: one toxin numbs the nervous system of insects, rendering them unable to sense or escape, while another toxin interferes with the autonomic nervous system of vertebrates to make small vertebrates easier to subdue and devour. The exact effects and makeup of the venom have not been thoroughly evaluated, in part because it is difficult to extract in significant quantities and quickly deteriorates when processed. ''S. heros'' bites are very painful to vertebrates. A rat bitten by ''S. heros'' in the leg showed signs of excruciating pain, followed by soreness, but returned to normal after five hours. For humans, a bite from ''S. heros'' usually causes sharp, searing, local pain and swelling, but has never caused any confirmed deaths. ''S. heros'' bites are known to occasionally cause
nausea Nausea is a diffuse sensation of unease and discomfort, sometimes perceived as an urge to vomit. While not painful, it can be a debilitating symptom if prolonged and has been described as placing discomfort on the chest, abdomen, or back of the ...
, headache, and localized skin necrosis. However, there are individual cases of severe symptoms and injury (including
kidney failure Kidney failure, also known as end-stage kidney disease, is a medical condition in which the kidneys can no longer adequately filter waste products from the blood, functioning at less than 15% of normal levels. Kidney failure is classified as eit ...
due to rhabdomyolysis, and
heart attack A myocardial infarction (MI), commonly known as a heart attack, occurs when blood flow decreases or stops to the coronary artery of the heart, causing damage to the heart muscle. The most common symptom is chest pain or discomfort which ma ...
) in humans resulting from ''Scolopendra'' bites.


References

{{Taxonbar, from=Q2637538 Scolopendridae Arthropods of Mexico Arthropods of the United States Fauna of the Southwestern United States Fauna of the Chihuahuan Desert Fauna of the Sonoran Desert Animals described in 1853 Taxa named by Charles Frédéric Girard