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The scarce large blue (''Phengaris teleius'') is a species of
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Lycaenidae Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfl ...
. It is found in
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
,
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
, the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
,
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
,
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
, the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
,
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...
,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
, northern
Serbia Serbia (, ; Serbian language, Serbian: , , ), officially the Republic of Serbia (Serbian language, Serbian: , , ), is a landlocked country in Southeast Europe, Southeastern and Central Europe, situated at the crossroads of the Pannonian Bas ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
,
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
, and
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
and East across the
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. The species was first described by
Johann Andreas Benignus Bergsträsser Johann Andreas Benignus Bergsträsser (21 December 1732, in Idstein – 24 December 1812, in Hanau) was a German educator, philologist, and entomologist. Education He studied philology and theology at the universities of Jena and Halle, and ...
in 1779.


Biology

The larva of this species first feeds on ''
Sanguisorba officinalis ''Sanguisorba officinalis'', commonly known as great burnet, is a plant in the family Rosaceae, subfamily Rosoideae. It is native throughout the cooler regions of the Northern Hemisphere in Europe, northern Asia, and northern North America. It ...
'' (great burnet), then moves onto ant nests and is a predator of the ant brood. ''
Myrmica rubra ''Myrmica rubra'', also known as the common red ant or erroneously the European fire ant, is a species of ant of the genus '' Myrmica'', found all over Europe and is now invasive in some parts of North America and Asia. It is mainly red in colou ...
'' and '' Myrmica scabrinodis'' have been reported as frequent host ant species. A recent microhabitat preference study indicates that grazing is necessary for maintaining the present distribution and abundance of these butterflies.


Description from Seitz

''L. euphemus'' Hbn. (= diomedes Rott.) (83 a), Male above almost entirely blue, but not shining; the margins, the discocellular spot of the forewing and commonly some small spots on the disc black; female much blacker, often a little paler on the disc, this lighter area bearing rows of black spots. Underside with very numerous ocelli, which are rarely as large and conspicuous as in our figure of the underside. The species is at once distinguished from the very similar ''
arion Arion (; grc-gre, Ἀρίων; fl. c. 700 BC) was a kitharode in ancient Greece, a Dionysiac poet credited with inventing the dithyramb. The islanders of Lesbos claimed him as their native son, but Arion found a patron in Periander, tyrant ...
'' by the underside not bearing an ocellus in the cell proximally to the discocellular spot. Throughout Europe (with the exception of England) and the adjacent districts of Asia, from North Germany and Russia to Italy, and from Paris to Dauria. — Near the North- Western boundary there occur especially small individuals (of the size of ''icarus''), which have but few ocelli; these are ab. ''paula'' Schultz. In ab. ''obsoleta'' Gillm. the ocelli of the hindwing are entirely absent or almost, while in ab. ''striata'' Gillm. they are modified into streaks. — ''obscurata'' Stgr. (83 b) is a strongly darkened form from Central Asia (also already in the Ural) — ''euphemia'' Stgr. (83 b) is much larger than European specimens, with a broad block border which sharply contrasts with the light blue; from Amurland. North China and Corea. — ''kazamoto'' Druce (83 b), from Japan, is above uniformly black-brown above in both sexes, without any trace of blue, and the underside is more strongly ocellate. — Egg semiglobular with the top sunk in, greenish white, laid on ''Sanguisorba'' (usually on the inflorescence). The young larva purplish brown with black head and pale segmental incisions; it bores into the heads of the plant and lives later on in the seed-pods; it hibernates. Pupa on the ground, under stones, clods of earth, and the dead leaves of the food-plant. The butterflies are sporadic, their localities being widely dispersed. They fly in damp meadows where ''Sanguisorba'' grows, being here usually very abundant. They settle almost exclusively on Sanguisorba: when disturbed they fly mostly only as far as the nearest cluster of that plant, where they settle on a flower with the wings always closed, their flight being rather slow and flapping. In July and August.Seitz, A. ed. Band 1: Abt. 1, ''Die Großschmetterlinge des palaearktischen Faunengebietes, Die palaearktischen Tagfalter'', 1909, 379 Seiten, mit 89 kolorierten Tafeln (3470 Figuren)


Subspecies

There are six
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. *''P. t. teleius'' central Europe,
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...
and western
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
*''P. t. chosensis'' (Matsumura, 1927) southern
Ussuri The Ussuri or Wusuli (russian: Уссури; ) is a river that runs through Khabarovsk and Primorsky Krais, Russia and the southeast region of Northeast China. It rises in the Sikhote-Alin mountain range, flowing north and forming part of the Si ...
*''P. t. euphemia'' (Staudinger, 1887)
Amur Oblast Amur Oblast ( rus, Аму́рская о́бласть, r=Amurskaya oblast, p=ɐˈmurskəjə ˈobləsʲtʲ) is a federal subject of Russia (an oblast), located on the banks of the Amur and Zeya Rivers in the Russian Far East. The administrative ...
, Ussuri *''P. t. obscurata'' (Staudinger, 1892)
Transbaikalia Transbaikal, Trans-Baikal, Transbaikalia ( rus, Забайка́лье, r=Zabaykalye, p=zəbɐjˈkalʲjɪ), or Dauria (, ''Dauriya'') is a mountainous region to the east of or "beyond" (trans-) Lake Baikal in Far Eastern Russia. The steppe and ...
*''P. t. sinalcon'' Murayama, 1992 northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
*''P. t. splendens'' (Kozhantshikov, 1924)
Altai mountains The Altai Mountains (), also spelled Altay Mountains, are a mountain range in Central Asia, Central and East Asia, where Russia, China, Mongolia and Kazakhstan converge, and where the rivers Irtysh and Ob River, Ob have their headwaters. The m ...
,
Sayan Mountains The Sayan Mountains (russian: Саяны ''Sajany''; mn, Соёны нуруу, ''Soyonï nurû''; otk, 𐰚𐰇𐰏𐰢𐰤, Kögmen) are a mountain range in southern Siberia, Russia (Buryatia, Irkutsk Oblast, Krasnoyarsk Krai, Tuva Republic ...


Sources

* * {{Taxonbar, from=Q1340609 Phengaris Butterflies of Europe Butterflies of Japan Butterflies described in 1779 Taxonomy articles created by Polbot