The Scalopinae, or New World moles, are one of three subfamilies of the family

Talpidae The family Talpidae () includes the moles (some of whom are called shrew moles and desmans) who are small insectivorous mammals of the order Eulipotyphla. Talpids are all digging animals to various degrees: moles are completely subterranean a ...

, which consists of moles and mole-like animals; the other two subfamilies being the Old World talpids (the

Talpinae The subfamily Talpinae, sometimes called "Old World moles" or "Old World moles and relatives", is one of three subfamilies of the mole family Talpidae, the others being the Scalopinae, or New World moles, and the Uropsilinae, or shrew-like moles. ...

) and the Chinese shrew-like moles (

Uropsilinae The shrew moles or shrew-like moles (''Uropsilus'') are shrew-like members of Talpidae, the mole family of mammals endemic to the forested, high-alpine region bordering China, Myanmar, and Vietnam. They possess a long snout, a long slender tail, ...

). The Scalopinae are the only Talpidae subfamily to consist entirely of undisputed

moles Moles can refer to: * Moles de Xert, a mountain range in the Baix Maestrat comarca, Valencian Community, Spain * The Moles (Australian band) *The Moles, alter ego of Scottish band Simon Dupree and the Big Sound People *Abraham Moles, French engin ...

and no mole-like close relatives such as

shrew-moles A shrew mole or shrew-mole is a Mole (animal), mole that resembles a shrew. Species with this name include: * Five species in the subfamily Uropsilinae, or "shrew-like moles", native to China: **Equivalent teeth shrew mole (''Uropsilus aequodonen ...

desmans The desman, a snouted and naked-tailed diving insectivore of the tribe Desmanini (also considered a subfamily, Desmaninae), belongs to one of two Eurasian species of the mole family, Talpidae. This tribe consists of two monotypic genera of semi ...

. They are found in temperate

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

and parts of

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

. In

, they exist virtually everywhere soil conditions permit except northern

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

and those areas of northeastern

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

where the soil is too sandy. Despite this subfamily being referred to as "New World moles", there are also two species in the mountains of

, each in their own

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

genus. Morphological and paleontological analyses indicate that both tribes in the subfamily originated in Eurasia during the

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

and migrated to North America during the Neogene, with the Condylurini later going extinct throughout all of their Eurasian range. The Scalopini also migrated at least two different times from North America back to Eurasia, with the two Chinese species likely originating from this. In addition, phylogenetic and morphological evidence supports the Condylurini not belonging in the Scalopinae, and occupying a much more basal position in

Taxonomy

The Scalopininae are divided into two tribes, six genera, and nine species: *Tribe

Condylurini ''Condylura'' is a genus of moles that contains a single extant species, the star-nosed mole ''(Condylura cristata)'' endemic to the northern parts of North America. It is also the only living member of the tribe Condylurini. While today endemic ...

** Genus ''

Condylura ''Condylura'' is a genus of moles that contains a single extant species, the star-nosed mole ''(Condylura cristata)'' endemic to the northern parts of North America. It is also the only living member of the tribe Condylurini. While today endemic ...

'', the star-nosed mole *Tribe

Scalopini The Scalopini are a tribe of moles belonging to the family Talpidae. They include all the New World moles apart from the strikingly distinctive star-nosed mole. As the similarity of the names implies, they are the standard form of the Scalopinae ...

** Genus ''

Scalopus The Eastern Mole or Common Mole (''Scalopus aquaticus'') is a medium-sized North American mole. It is the only member of genus ''Scalopus''. It is found in forested and open areas with moist sandy soils in northern Mexico, the eastern Unite ...

'', the common eastern mole ** Genus ''

Scapanus ''Scapanus'' is a genus of mole (animal), moles in the family Talpidae. They live in North America from west of the Rocky Mountains, Rockies south to Baja California, Baja California del Norte, and north to British Columbia, wherever conditions ...

'', five species of western moles ** Genus ''

Parascalops The hairy-tailed mole (''Parascalops breweri''), also known as Brewer's mole, is a medium-sized North American mole. It is the only member of the genus ''Parascalops''. The species epithet ''breweri'' refers to Thomas Mayo Brewer, an American n ...

'', the hairy-tailed mole ** Genus ''

Scapanulus The Gansu mole (''Scapanulus oweni'') is a species of mammal in the family Talpidae endemic to central China, where it occurs in Shaanxi, Gansu, Sichuan, and Qinghai. It is the only species in the genus ''Scapanulus''. Along with the Medog ...

'', the Gansu mole **Genus '' Alpiscaptulus'', the Medog mole

References

Talpidae {{Eulipotyphla-stub