''Sarracenia flava'', the yellow pitcherplant, is a

carnivorous plant Carnivorous plants are plants that derive some or most of their nutrients from trapping and consuming animals or protozoans Protozoa (singular: protozoan or protozoon; alternative plural: protozoans) are a group of single-celled eukaryot ...

in the family

Sarraceniaceae Sarraceniaceae are a family of pitcher plants, belonging to order Ericales (previously Nepenthales). The family comprises three extant genera: ''Sarracenia'' (North American pitcher plants), '' Darlingtonia'' (the cobra lily or California pitch ...

. Like all the Sarraceniaceae, it is native to the

New World The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. 3 ...

. Its range extends from southern

Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...

, through

Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...

and

Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...

, to the coastal plains of southern

Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...

North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...

and

South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...

. Populations also exist in the

Piedmont it, Piemontese , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1_info1 = , demographics1_title2 ...

and mountains of

. Like other members of the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

Sarracenia ''Sarracenia'' ( or ) is a genus comprising 8 to 11 species of North American pitcher plants, commonly called trumpet pitchers. The genus belongs to the family Sarraceniaceae, which also contain the closely allied genera '' Darlingtonia'' and ...

'', the yellow pitcher plant traps insects using a rolled leaf, which in this species is a vibrant yellow in color, and up to over a meter (3 ft) in heightGraham, D.L. 1997. ''

Carnivorous Plant Newsletter The ''Carnivorous Plant Newsletter'' is the official publication of the International Carnivorous Plant Society (ICPS), the largest such organization in the world. It is headquartered in Walnut Creek, California. History and editorship The newsle ...

'' 26(4): 118–120. (although 50 cm, 20" is more typical). The uppermost part of the leaf is flared into a lid (the operculum), which prevents excess rain from entering the pitcher and diluting the digestive secretions within. The upper regions of the pitcher are covered in short, stiff, downwards-pointing hairs, which serve to guide

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

s alighting on the upper portions of the leaf towards the opening of the pitcher tube. The upper regions are also brightly patterned with flower-like

anthocyanin Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical compo ...

markings, particularly in the varieties ''S. flava'' var. ''rugelii'' and ''S. flava'' var. ''ornata'': these markings also serve to attract insect prey. The opening of the pitcher tube is retroflexed into a 'nectar roll' or

peristome Peristome (from the Greek ''peri'', meaning 'around' or 'about', and ''stoma'', 'mouth') is an anatomical feature that surrounds an opening to an organ or structure. Some plants, fungi, and shelled gastropods have peristomes. In mosses In mosses, ...

, whose surface is studded with

nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...

-secreting glands. The nectar contains not only

sugar Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose, fructose, and galactose. Compound sugars, also called disaccharides or double ...

s, but also the

alkaloid Alkaloids are a class of basic, naturally occurring organic compounds that contain at least one nitrogen atom. This group also includes some related compounds with neutral and even weakly acidic properties. Some synthetic compounds of similar ...

coniine Coniine is a poisonous chemical compound, an alkaloid present in and isolable from poison hemlock ('' Conium maculatum''), where its presence has been a source of significant economic, medical, and historico-cultural interest; coniine is also prod ...

toxin A toxin is a naturally occurring organic poison produced by metabolic activities of living cells or organisms. Toxins occur especially as a protein or conjugated protein. The term toxin was first used by organic chemist Ludwig Brieger (1849– ...

also found in hemlock), which probably intoxicates the prey. Prey entering the tube find that their footing is made extremely uncertain by the smooth,

wax Waxes are a diverse class of organic compounds that are lipophilic, malleable solids near ambient temperatures. They include higher alkanes and lipids, typically with melting points above about 40 °C (104 °F), melting to give low ...

y secretions found on the surfaces of the upper portion of the tube. Insects losing their footing on this surface plummet to the bottom of the tube, where a combination of digestive fluid, wetting agents and inward-pointing hairs prevent their escape. Some large insects (such as

wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...

s) have been reported to escape from the pitchers on occasion, by chewing their way out through the wall of the tube. Red Sarracenia flava

spring Spring(s) may refer to: Common uses * Spring (season), a season of the year * Spring (device), a mechanical device that stores energy * Spring (hydrology), a natural source of water * Spring (mathematics), a geometric surface in the shape of a ...

, the plant produces large

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

s with 5-fold

symmetry Symmetry (from grc, συμμετρία "agreement in dimensions, due proportion, arrangement") in everyday language refers to a sense of harmonious and beautiful proportion and balance. In mathematics, "symmetry" has a more precise definit ...

. The yellow

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

s are long and strap-like, and dangle over the

umbrella An umbrella or parasol is a folding canopy supported by wooden or metal ribs that is usually mounted on a wooden, metal, or plastic pole. It is designed to protect a person against rain or sunlight. The term ''umbrella'' is traditionally used ...

-like style of the flower, which is held upside down at the end of a 50 cm, 20" long scape. The stigma of the flower are found at the tips of the 'spokes' of this umbrella. Pollinating insects generally enter the flower from above, forcing their way into the cavity between the petals and umbrella, and depositing any pollen they are carrying on the stigmata as they enter. The pollinators generally exit the flower, having been dusted with the plant's own pollen, by lifting a petal. This one-way system helps to ensure

cross pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, a ...

. In late summer and autumn, the plant stops producing carnivorous leaves, and instead produces flat, non-carnivorous phyllodia. This is probably an adaptation to low light levels and insect scarcity during the winter months, and shows clearly the cost of carnivory. The yellow pitcher plant is easy to cultivate, and is one of the most popular carnivorous plants in

horticulture Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...

. The yellow pitcher plant readily hybridises with other members of the genus ''Sarracenia'': the hybrids ''S. x catesbaei'' (''S. flava'' × ''S. purpurea'') and ''S. moorei'' (''S. flava'' × ''S. leucophylla'') are found in the wild, and are also popular amongst collectors. Image:Sarracenia flavaPCCA20060423-3480B.jpg, Pitcher mouth with operculum Image:Sarracenia flavaPCCA20060423-3391B.jpg, Flower Image:SarraceniaFlavaVars.JPG, A "pitcherplant meadow" in the Florida panhandle, with mixed varieties of ''Sarracenia flava'': var. ''rugelii'', var. ''ornata'', and var. ''rubricorpora''

References

* *Mody, NV; Henson, R; Hedin, PA; Kokpol, U; Miles, DH (1976). Isolation of insect paralyzing agent coniine from ''Sarracenia flava''. ''Cellular and Molecular Life Sciences'' 32(7): 829–830.
N.C. State University reference
*

External links

Botanical Society of America, ''Sarracenia'' - the Pitcher Plants
{{Taxonbar, from=Q1499289 flava Carnivorous plants of North America Endemic flora of the United States Flora of the Southeastern United States Least concern flora of the United States Plants described in 1753 Taxa named by Carl Linnaeus