The Erycinae, known as the Old World sand boas, are a subfamily of nonvenomous snakes in the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Boidae The Boidae, commonly known as boas or boids, are a family of nonvenomous snakes primarily found in the Americas, as well as Africa, Europe, Asia, and some Pacific Islands. Boas include some of the world's largest snakes, with the green anacond ...

Species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of the subfamily Erycinae are found in Europe,

Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...

, Africa,

Arabia The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...

, central and southwestern Asia, India, Sri Lanka, and western North America. Four genera comprising 18

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

are currently recognized as being valid.

Description

Erycinae is a subfamily of stout-bodied snakes, all of which are competent burrowers. The largest, '' E. johnii'', rarely exceeds in total length (including tail). Most grow to around in total length. They have small eyes and hard, small scales to protect their skin from the grit of sand. A great deal of

sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...

exists, with females generally becoming much larger than males. Erycines have skeletal adaptations to burrowing. The skull is more compact than in the subfamily Boinae. Also, the vertebrae of the tail are increased in size but reduced in number.

Distribution and habitat

Erycines are found in Southeastern Europe, Asia Minor, North, Central, West and East Africa, Arabia, Central and Southwest Asia, India, Sri Lanka, southwestern Canada, the western United States, and northwestern Mexico. McDiarmid RW, Campbell JA, Touré T (1999). ''Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1''. Washington, District of Columbia: Herpetologists' League. 511 pp. (series). (volume). Fossil erycines have been found in rock strata over 50 million years old, and were once widespread in North America. Now, only four

remain in North America, as well as the sand boas in Africa, Asia, and Southeastern

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...

. The oldest known erycines are from the

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...

-aged

Messel Pit The Messel pit (german: Grube Messel) is a disused quarry near the village of Messel (Landkreis Darmstadt-Dieburg, Hesse) about southeast of Frankfurt am Main, Germany. Bituminous shale was mined there. Because of its abundance of well-preserved ...

in Germany.

Behavior

The majority of sand boas spend much of their time basking below the surface of the sand, with only their eyes or head exposed. When potential prey approaches, they erupt out of the sand, bite, and employ

constriction Constriction is a method used by various snake species to kill or subdue their prey. Although some species of venomous and mildly venomous snakes do use constriction to subdue their prey, most snakes which use constriction lack venom. The snake i ...

to subdue it.

Feeding

The erycines' primary diet is rodents, but they have also been known to prey on lizards and birds.

Reproduction

Otherwise far removed from their boine relatives, erycines are generally

ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...

, i.e., giving birth to live young. At least three species lay eggs, however: the

Calabar python :''Common names: Calabar ground boa, burrowing boa, Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . Calabar boa, more.'' The Calabar python (''Calabaria reinhardtii'') is a species of non-ven ...

, ''Calabaria reinhardtii'' (once regarded as a

python Python may refer to: Snakes * Pythonidae, a family of nonvenomous snakes found in Africa, Asia, and Australia ** ''Python'' (genus), a genus of Pythonidae found in Africa and Asia * Python (mythology), a mythical serpent Computing * Python (pro ...

for this reason), the Arabian sand boa, '' Eryx jayakari'', and the West African sand boa, ''E. muelleri''.

Smuggling and poaching in India

Poaching and smuggling of sand boas is often reported in India. Most of the smuggled snakes go to the United States. There is a misconception about their medicinal and aphrodisiacal properties, as well as the belief that keeping this snake as a pet brings wealth and prosperity.

Captivity

Eryx colubrinus ''Eryx colubrinus'', the Egyptian or Kenyan sand boa,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . is a species of snake in the family Boidae. The species is endemic to Northern and Eastern ...

'', '' E. conicus'' and '' E. johnii'' are frequently available in the exotic pet trade and are often captive-bred. They breed readily, their small size making them an attractive option. They are usually not aggressive species, though they sometimes have a tendency to bite, and also spend the vast majority of their time hiding. Other species are not commonly available, but are occasionally imported. Sand Boa

Genera

* Not including the nominate subspecies.
^T

Type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal ...

References

External links

The Sand Boa Page
a
Kingsnake.com
Accessed 16 July 2008. {{Taxonbar, from=Q2220578 Tetrapod subfamilies Taxa named by Charles Lucien Bonaparte