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The South American land mammal ages (SALMA) establish a
geologic timescale The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronol ...
for
prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
n
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
beginning 64.5 Ma during the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
and continuing through to the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...
(0.011 Ma). These periods are referred to as ages, stages, or intervals and were established using geographic place names where
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
materials where obtained.Flynn & Swisher, 1995 The basic unit of measure is the first/last boundary statement. This shows that the first appearance event of one
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
is known to predate the last appearance event of another. If two taxa are found in the same fossil
quarry A quarry is a type of open-pit mine in which dimension stone, rock, construction aggregate, riprap, sand, gravel, or slate is excavated from the ground. The operation of quarries is regulated in some jurisdictions to reduce their envi ...
or at the same stratigraphic horizon, then their age-range zones overlap.


Background

South America was an island continent for much of the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
, or the "Age of Mammals". As a result, its mammals evolved in their own unique directions, as Australia and Madagascar still have today.


Paleogeographic timeline

A simplified
paleogeographic Palaeogeography (or paleogeography) is the study of historical geography, generally physical landscapes. Palaeogeography can also include the study of human or cultural environments. When the focus is specifically on landforms, the term paleo ...
timeline of South America:Darin A. Croft - South American Fossil Mammals
/ref> * 66 Ma – South America was connected to both North America and Antarctica. Soon after this point, it lost its connection to North America. * 66–50 Ma –
Tiupampan The Tiupampan ( es, Tiupampense) age is a period of geologic time (64.5–62.5 Ma) within the Paleocene epoch of the Paleogene used more specifically with South American land mammal ages (SALMA). It is the oldest SALMA age and precedes the Peligra ...
to
Casamayoran The Casamayoran ( es, Casamayorense) age is a period of geologic time (50.0–48.0 Ma) within the Early Eocene epoch of the Paleogene, used more specifically within the South American land mammal age (SALMA) classification. It follows the Itaboraia ...
- South America was connected to Antarctica which, in turn, was connected to Australia. The Antarctica–Australia connection was lost around the end of this interval or perhaps as much as 15 million years later. * 50–34 Ma – Casamayoran to Tinguirirican - South America was connected to Antarctica, which was not yet covered by ice. * 34 Ma –
Tinguirirican The Tinguirirican ( es, Tinguiririquense) age is a period of geologic time (36.0–29.0 Ma) within the Late Eocene and Early Oligocene epochs of the Paleogene, used more specifically within the SALMA classification in South America. It follows th ...
- South America and Antarctica became detached and glaciations started to form in Antarctica. * 34–9 Ma – Tinguirirican to
Chasicoan The Chasicoan ( es, Chasiquense or es, Chasicoense) age is a period of geologic time from 10–9 Ma within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Mayoan and ...
- South America had no land connections to any other continent. * 9–3 Ma –
Huayquerian The Huayquerian ( es, Huayqueriense) age is a period of geologic time (9.0–6.8 Ma) within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification. It follows the Mayoan and precedes the Montehermosan age. ...
to
Chapadmalalan The Chapadmalalan age is a period of geologic time (4.0–3.0 Ma) within the Pliocene epoch of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan and precedes the Uquian age. Fossil content F ...
- islands formed between South and North America. A complete Isthmus of Panama most likely formed near the end of this interval, leading to the
Great American Biotic Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
(GABI). * 3 Ma to present –
Uquian The Uquian age is a period of geologic time (3.0–1.5 Ma) within the Pliocene and Pleistocene epochs of the Neogene used more specifically with South American Land Mammal Ages. It follows the Montehermosan and precedes the Ensenadan The Ensenad ...
to
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
- the land connection between South and North America is established and migration between the formerly separated continents occurs. The main migrational route is from north to south, rather than the opposite way. This led to much higher extinction levels of groups in South America than in North America. * Pleistocene – the
glacial A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate betw ...
s and
interglacial An interglacial period (or alternatively interglacial, interglaciation) is a geological interval of warmer global average temperature lasting thousands of years that separates consecutive glacial periods within an ice age. The current Holocene in ...
s of the Pleistocene caused drastic
eustatic sea level The eustatic sea level is the distance from the center of the earth to the sea surface. An increase of the eustatic sea level can be generated by decreasing glaciation, increasing spreading rates of the mid-ocean ridges or more mid-oceanic ridges. ...
changes, widening and narrowing the land bridge at the 'bottleneck' of Panama. As a side-effect, the vegetation changed during this period of strong climatic changes. * Late Pleistocene – the earliest humans arrived in South America and settled in various parts of the continent. Evidence for cohabitation with the latest
Pleistocene megafauna Pleistocene megafauna is the set of large animals that lived on Earth during the Pleistocene epoch. Pleistocene megafauna became extinct during the Quaternary extinction event resulting in substantial changes to ecosystems globally. The role of hu ...
has been found at multiple locations, such as
Monte Verde Monte Verde is an archaeological site in the Llanquihue Province in southern Chile, located near Puerto Montt, Southern Chile, which has been dated to as early as 18,500 cal BP (16,500 BC). Previously, the widely accepted date for early occup ...
in coastal
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
and
Tibitó Tibitó is the second-oldest dated archaeological site on the Altiplano Cundiboyacense, Colombia.Altiplano Cundiboyacense The Altiplano Cundiboyacense () is a high plateau located in the Eastern Cordillera of the Colombian Andes covering parts of the departments of Cundinamarca and Boyacá. The altiplano corresponds to the ancient territory of the Muisca. The Alti ...
in
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
.


Definitions


Cenozoic fossiliferous stratigraphic units in South America

The following formations have provided vertebrate, insect or plant fossils, formations with other invertebrates are excluded:


Fossil content


See also

*
Biochronology In paleontology, biochronology is the correlation in time of biological events using fossils. In its strict sense, it refers to the use of assemblages of fossils that are not tied to stratigraphic sections (in contrast to biostratigraphy, where th ...
**
North American land mammal age The North American land mammal ages (NALMA) establishes a geologic timescale for North American fauna beginning during the Late Cretaceous and continuing through to the present. These periods are referred to as ages or intervals (or stages when ref ...
(NALMA) **
Asian land mammal age The Asian land mammal ages, acronym ALMA, establish a geologic timescale for prehistoric Asian fauna beginning 58.7 Mya during the Paleogene and continuing through to the Miocene ( Aquitanian) (23.03 Ma). These periods are referred to as ages, s ...
(ALMA) **
European land mammal age The European Land Mammal Mega Zones (abbreviation: ELMMZ, more commonly known as European land mammal ages or ELMA) are zones in rock layers that have a specific assemblage of fossils (biozones) based on occurrences of fossil assemblages of Europe ...
(ELMA or ELMMZ) *
List of fossiliferous stratigraphic units in Colombia Several stratigraphic units in Colombia have provided fossils. The richest formations are the Devonian Cuche and Floresta Formations, the Cretaceous Paja Formation, the Paleocene Cerrejón Formation and the Miocene La Venta site. The latter is ...
*
List of fossiliferous stratigraphic units in Ecuador This is a list of fossiliferous stratigraphic units in Ecuador. __NOTOC__ List of fossiliferous stratigraphic units See also * * List of gomphothere fossils in South America#Ecuador, Gomphothere fossils in Ecuador * List of fossiliferous s ...
*
List of fossiliferous stratigraphic units in Paraguay This is a list of fossiliferous stratigraphic units in Paraguay. __NOTOC__ List of fossiliferous stratigraphic units See also * * South American land mammal age * List of gomphothere fossils in South America#Paraguay, Gomphothere fossil ...
*
List of fossiliferous stratigraphic units in Uruguay This is a list of fossiliferous stratigraphic units in Uruguay. __NOTOC__ List of fossiliferous stratigraphic units See also * * South American land mammal ages * List of gomphothere fossils in South America#Uruguay, Gomphothere fossils i ...
*
List of fossiliferous stratigraphic units in Venezuela This is a list of fossiliferous stratigraphic units in Venezuela. List of fossiliferous stratigraphic units See also * * List of gomphothere fossils in South America#Venezuela, Gomphothere fossils in Venezuela * List of fossiliferous str ...


Notes and references


Notes


References


Bibliography


* ;Paleogene * *


* * *


;Paleogene * ;Pleistocene * ;Austral Basin * * ;Santa María-Hualfín Basin * * ;Santa Fe Province * ;Aconquija Formation * * * ;Agua de la Piedra Formation * * * * * ;Aisol Formation * * ;Andalhuala Formation * ;Andesitas Huancache Formation * * ;Arroyo Chasicó Formation * * * ;Belgrano Formation * ;Bororó Formation * ;Brochero Formation * * ;Casamayor Formation * * ;Cerro Azul Formation * * * ;Cerro Bandera Formation * * ;Chapadmalal Formation * * * * ;Chichinales Formation * * ;Chiquimil Formation * * * * * ;Collón Curá Formation * * * * ;Deseado Formation * * ;Divisadero Largo Formation * ;La Ensenada Formation * * * ;Las Flores Formation, Sierra del Tontal * ;Las Flores Formation, Golfo San Jorge Basin * * * ;Fortín Tres Pozos Formation * * ;Gaiman Formation * * ;Geste Formation * * * ;Gran Bajo del Gualicho Formation * ;Hernandarias Formation * ;Huayquerías Formation * * ;Huitrera Formation * ;India Muerta Formation * ;La Invernada Formation * ;Ituzaingó Formation * * * ;Koluel Kaike Formation * ;Laguna Brava Formation * ;Laguna del Hunco Formation * ;Lefipán Formation * ;Leticia Formation * ;Loma de Las Tapias Formation * * * ;Luján Formation * * ;Lumbrera Formation * * * ;Maimará Formation * * * ;Maíz Gordo Formation * ;Mariño Formation * * ;Mealla Formation * ;Miramar & San Andrés Formations * * * ;Monte Hermoso Formation * * * * * ;Monte León Formation * ;El Morterito Formation * ;Palo Pintado Formation * * * ;Paraná Formation * * * ;Peñas Coloradas Formation * * ;Pinturas Formation * * ;Piquete Formation * ;Playa de Los Lobos Allo Formation * ;Puerta del Diablo Formation * ;Puerto Madryn Formation * * ;Quebrada de Los Colorados Formation * * * * ;Río Chico Group * ;Río Foyel Formation * * ;Río Loro Formation * ;Río Mayo Formation * ;Río Negro Formation * ;Río Quinto Formation * ;Saladillo Formation * ;Salamanca Formation * * * * * ;Saldungaray Formation * ;Salicas Formation * ;Santa Cruz Formation * * * * * * * * * * * * * * * * ;Sarmiento Formation & Colhué Huapí Member * * * * * * * * * * ;Toro Negro Formation * ;Uquía Formation * * * ;Vaca Mahuida Formation * ;Ventana Formation * * ;Vorohué Formation * * * ;Yupoí Formation *


* * * ;Casira Formation * ;Cerdas beds * ;Honda Group * * * * ;Lacayani fauna * ;Ñuapua Formation * ;Quehua Formation * ;Salla Formation * * * ;Santa Lucía Formation * * * * * * * * * ;Tarija Formation * ;Umala Formation * ;Yecua & Petaca Formations *


* ;Entre-Corrégos Formation * ;Fonseca Formation * ;Graxaim Formation * ;Guabirotuba Formation * ;Içá Formation * ;Itaboraí Formation * * * * * * ;Maria Farinha Formation * ;Pirabas Formation * ;Rio Madeira Formation * ;Santa Vitória Formation * ;Solimões Formation * * * * * ;Touro Passo Formation * ;Tremembé Formation * * *


;Abanico Formation * * * * * * ;Bahía Inglesa Formation * * * * * * * * * * * * * ;Caleta Herradura Formation * ;Chíu-Chíu Formation * ;Chucal Formation * ;Coquimbo Formation * ;Cura-Mallín Group * * * ;Horcón Formation * ;Huaylas Formation * * ;Loreto Formation * * * * ;Navidad Formation * ;La Portada Formation * ;Río Baguales Formation * ;Río Frías Formation * * * * *


;Altiplano Cundiboyacense * * * * * * * * * * * * ;Cesar-Ranchería Basin * * * * * * * * * * * * * ;Cocinetas Basin * * * * * * * * * ;Barzalosa Formation * ;Gualanday Group * ;Honda Group * * * * * * * * * * * * * * * * * * * * * * * * * * * * ;Los Hoyos beds * ;Mugrosa Formation * ;Rotinet Formation * ;Sincelejo Formation *


;Biblián Formation * ;Dos Bocas Formation * * ;Letrero Formation * ;Onzole Formation * ;Seca Formation *


;Chaco Formation * *


;Peruvian Amazon * * ;Sechura Basin * ;Chambira Formation * * * ;Chilcatay Formation * * * * ;Chota Formation * ;Iñapari Formation * * ;Madre de Dios Formation * * * ;Mogollón Formation * * ;Moquegua Formation * * ;Muñani Formation * ;Paracas Formation * ;Pisco Formation * * * * * * * * * * * * * * ;Pozo Formation * * ;Soncco Formation * ;Yahuarango Formation * * * ;Yumaque Formation *


;Pleistocene * ;Camacho Formation * * * ;Dolores Formation * * * ;Fray Bentos Formation * * ;Raigón Formation * ;San José Formation * ;Sopas Formation *


;Chaguaramas Formation * ;Parángula Formation * ;Falcón Basin ;Cantaure & Paraguaná Formations * ;Capadare Formation * * ;Castillo Formation * * * * * ;Mesa Formation * ;Río Yuca Formation * ;San Gregorio Formation * * ;Santa Inés Formation * ;Urumaco, Socorro & Codore Formations * * * * *


Further reading

* * * * * * * * * * * * * {{SALMA * South America