Salix Glauca
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''Salix glauca'' is a species of flowering plant in the willow family known by the common names gray willow, grayleaf willow, white willow, and glaucous willow. It is native to North America, where it occurs throughout much of
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
, northern and western Canada, and the contiguous United States south through the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
to northern
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
.Uchytil, Ronald J. 1992
''Salix glauca''.
In: Fire Effects Information System, nline U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory.
It can also be found in
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
, northwestern Europe, and Siberia.


Description

This
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...
is usually a
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
growing up to tall, but in appropriate habitat it becomes a tree up to tall. The smooth gray bark becomes furrowed with age. The species is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
, with male and female reproductive parts occurring on separate individuals. This species has secondary
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
, with male and female individuals different in function or morphology in aspects other than their reproductive structures. For example, female plants are more sensitive to
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
conditions. The seed stays on the plant until fall, when it is dispersed. The seed is coated in downy fibers that help it disperse on the wind and on water. Unlike the seeds of many other willows, these do not
germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
immediately on contact with the substrate, but overwinter under the snow and sprout in the spring. This provides cold
stratification Stratification may refer to: Mathematics * Stratification (mathematics), any consistent assignment of numbers to predicate symbols * Data stratification in statistics Earth sciences * Stable and unstable stratification * Stratification, or st ...
to the seeds, and allows them a few weeks more to develop than in summer-dispersing willows.


Distribution and habitat

In the northern part of its range, this plant codominates with other species of willow on
floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...
s and in shrubby riparian and
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
habitat. It may also grow scattered throughout coniferous forests and woodlands, dominated often by
spruce A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
s. In the southern part of its range, it grows in
alpine Alpine may refer to any mountainous region. It may also refer to: Places Europe * Alps, a European mountain range ** Alpine states, which overlap with the European range Australia * Alpine, New South Wales, a Northern Village * Alpine National Pa ...
and
subalpine climates Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
. Like many other willows, it colonizes freshly cleared habitat, such as floodplains recently scoured by water and forests recently burned.


Taxonomy

The taxonomy of ''S. glauca'' has been described as "confusing". With considerable geographic variation across its wide circumboreal-polar range, ''S. glauca'' may be considered "a very widespread and polymorphic species or species group", with currently no consensus whether it should be subdivided into races, subspecies or varieties. Formally and informally, there are a number of recognized subspecies (such as ''glauca'', ''stipulifera'', ''acutifolia'', ''callicarpaea'') and varieties (such as ''acutifolia'', ''glauca'', ''stipulata'', ''villosa''), but there are only small morphological differences to tell them apart. Furthermore, ''S. glauca'' is known to form hybrids with other willows, resulting in intermediates that are visually difficult to distinguish from one another. Some varieties and subspecies have very specific or limited distribution, though. The hybrid ''S. arctophila × S. glauca'' subsp. ''callicarpaea'', for instance, is not found in Canada, and is common in eastern parts of
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
, but absent from the west, whereas ''S. glauca'' subsp. ''glauca'' is not found on Greenland at all.


Ecology

As with other willows, ''S. glauca'' is an important food source for a variety of animals, particularly wintering
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
s, providing them with a rich source of calcium and phosphorus. It is considered moderately important as moose
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
, and during the winter it constitutes much of the diet for
snowshoe hare The snowshoe hare (''Lepus americanus''), also called the varying hare or snowshoe rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet. The animal's feet prevent it from sink ...
s.


Use

Native Americans used parts of willows, including this species, for medicinal purposes, basket weaving, to make bows and arrows, and for building animal traps.


References


External links


The Nature ConservancyFlora of North America
{{Taxonbar, from=Q158458 glauca Plants described in 1753 Taxa named by Carl Linnaeus