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The saker falcon (''Falco cherrug'') is a large species of
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
. This species breeds from central Europe eastwards across the
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
to
Manchuria Manchuria is an exonym (derived from the endo demonym " Manchu") for a historical and geographic region in Northeast Asia encompassing the entirety of present-day Northeast China (Inner Manchuria) and parts of the Russian Far East (Outer Manc ...
. It is mainly migratory except in the southernmost parts of its range, wintering in
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, the
Arabian peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
, northern
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
and western
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. The saker falcon is the national bird of Hungary, the United Arab Emirates, and Mongolia.


Taxonomy and systematics

This
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
belongs to the close-knit hierofalcon complex. In this group, there is ample evidence for rampant hybridization and
incomplete lineage sorting Incomplete lineage sorting, also termed hemiplasy, deep coalescence, retention of ancestral polymorphism, or trans-species polymorphism, describes a phenomenon in population genetics when ancestral gene copies fail to coalesce (looking backwards i ...
which confounds analyses of
DNA sequence DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
data to a massive extent; molecular studies with small
sample size Sample size determination is the act of choosing the number of observations or Replication (statistics), replicates to include in a statistical sample. The sample size is an important feature of any empirical study in which the goal is to make stat ...
s can simply not be expected to yield reliable conclusions in the entire hierofalcon group. The
radiation In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or through a material medium. This includes: * ''electromagnetic radiation'', such as radio waves, microwaves, infrared, visi ...
of the entire living diversity of hierofalcons seems to have taken place in the
Eemian interglacial The Eemian (also called the last interglacial, Sangamonian Stage, Ipswichian, Mikulin, Kaydaky, penultimate,NOAA - Penultimate Interglacial Period http://www.ncdc.noaa.gov/global-warming/penultimate-interglacial-period Valdivia or Riss-Würm) wa ...
at the start of the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...
, a mere 130,000–115,000 years ago; the saker falcon represents a lineage that expanded out of northeastern Africa into the interior of southeastern Europe and Asia, by way of the eastern
Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...
region. In captivity, lanners and sakers can interbreed, and
gyrfalcon The gyrfalcon ( or ) (), the largest of the falcon species, is a bird of prey. The abbreviation gyr is also used. It breeds on Arctic coasts and tundra, and the islands of northern North America and the Eurosiberian region. It is mainly a reside ...
-saker hybrids are also available (see bird flu experiment described in "Ecology and status"). The specific name, ''cherrug'', comes from the Sindhi name ''charg'' for a female saker. The
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
''saker'' comes from the () meaning "falcon". Saker falcons at the northeast edge of the range in the Altai Mountains are slightly larger, and darker and more heavily spotted on the underparts than other populations. These, known as the
Altai falcon The Altai falcon (''Falco cherrug altaicus''?) is a large falcon of questionable taxonomic position. It is often considered to be a subspecies of the saker falcon (''Falco cherrug''). It used to have a high reputation among Central Asian falco ...
, have been treated in the past either as a distinct species ''"Falco altaicus"'' or as a
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
between saker falcon and gyrfalcon, but modern opinion (e.g. ) is to tentatively treat it as a form of saker falcon, until comprehensive studies of its
population genetics Population genetics is a subfield of genetics that deals with genetic differences within and between populations, and is a part of evolutionary biology. Studies in this branch of biology examine such phenomena as adaptation, speciation, and pop ...
and
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
are available.


Description

The saker falcon is a large hierofalcon, larger than the
lanner falcon The lanner falcon (''Falco biarmicus'') is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A l ...
and almost as large as
gyrfalcon The gyrfalcon ( or ) (), the largest of the falcon species, is a bird of prey. The abbreviation gyr is also used. It breeds on Arctic coasts and tundra, and the islands of northern North America and the Eurosiberian region. It is mainly a reside ...
at length with a wingspan of . Males weigh between and females . It resembles a larger but browner gyrfalcon. It is larger and more heavily built than the related
lanner falcon The lanner falcon (''Falco biarmicus'') is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A l ...
.Orta, J., Boesman, P., Sharpe, C.J. & Marks, J.S. (2019). Saker Falcon (''Falco cherrug''). In: del Hoyo, J., Elliott, A., Sargatal, J., Christie, D.A. & de Juana, E. (eds.). Handbook of the Birds of the World Alive. Lynx Edicions, Barcelona. (retrieved from https://www.hbw.com/node/53244 on 15 February 2019). Saker falcons tend to have variable plumage.
Male Male (symbol: ♂) is the sex of an organism that produces the gamete (sex cell) known as sperm, which fuses with the larger female gamete, or ovum, in the process of fertilization. A male organism cannot reproduce sexually without access to ...
s and
female Female (Venus symbol, symbol: ♀) is the sex of an organism that produces the large non-motile ovum, ova (egg cells), the type of gamete (sex cell) that fuses with the Sperm, male gamete during sexual reproduction. A female has larger gamet ...
s are similar, except in size, as are young birds, although these tend to be darker and more heavily streaked. The call is a sharp ''kiy-ee'' or a repeated ''kyak-kyak-kyak''.


Ecology

The saker falcon is a raptor of open grasslands preferably with some trees or cliffs. It often hunts by horizontal pursuit, rather than the peregrine's stoop from a height, and feeds mainly on
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s and birds. In Europe,
ground squirrel Ground squirrels are members of the squirrel family of rodents ( Sciuridae), which generally live on or in the ground, rather than trees. The term is most often used for the medium-sized ground squirrels, as the larger ones are more commonly known ...
s and
feral pigeon Feral pigeons (''Columba livia domestica'' or ''Columba livia forma urbana''), also called city doves, city pigeons, or street pigeons,Nagy, Kelsi, and Johnson, Phillip David. ''Trash animals: how we live with natures filthy, feral, invasive, an ...
s are the most common prey items. This species usually builds no nest of own, but lays its 3–6 eggs in an old stick nest in a tree which was previously used by other birds such as storks, ravens or buzzards. It also often nests on cliffs. Saker nests support a species-rich assemblage of
commensal Commensalism is a long-term biological interaction (symbiosis) in which members of one species gain benefits while those of the other species neither benefit nor are harmed. This is in contrast with mutualism, in which both organisms benefit fro ...
insects.


Status and conservation

BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
categorises this bird as
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
, due to a rapid population decline, particularly on the central Asian breeding grounds. Ever since the collapse of the
Soviet Union The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, it was nominally a federal union of fifteen national ...
, the
United Arab Emirates The United Arab Emirates (UAE; ar, اَلْإِمَارَات الْعَرَبِيَة الْمُتَحِدَة ), or simply the Emirates ( ar, الِْإمَارَات ), is a country in Western Asia (The Middle East). It is located at th ...
have been the main destination for thousands of falcons caught and sold illegally for hefty sums at the black market.
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
is estimated to lose up to 1,000 saker falcons per year. The species also faces pressure from habitat loss and destruction. The population was estimated to be between 7,200 and 8,800 mature individuals in 2004. However, sakers live at low densities across large ranges in remote regions, making distribution status difficult to assess. A climatic niche modelling study pinpointed certain remote areas for targeted population surveys. In the United States, Canada and Europe there are several captive breeding projects. The most dramatic decline of the saker falcon in Asia has been in
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
and
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
. In contrast, a strongly protected and relatively abundant population persists in
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
. Saker falcons are known to be very susceptible to
avian influenza Avian influenza, known informally as avian flu or bird flu, is a variety of influenza caused by viruses adapted to birds.
, individuals having been found infected with highly pathogenic H5N1 (in
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the A ...
) and H7N7 (in
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
) strains. Therefore, an experiment was done with
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
gyr-saker falcons, which found that five falcons vaccinated with a commercial H5N2 influenza vaccine survived infection with a highly pathogenic H5N1 strain, whereas five unvaccinated falcons died. This means that sakers could be protected from bird flu by vaccination, at least in captivity. Faucon sacre MHNT.jpg, Egg Saker Falcon RWD3.jpg, Captive, Carolina Raptor Center, United States Saker_Falcon_RWD4.jpg, Captive, Carolina Raptor Center Sakerfalcon.JPG, Captive bird, Chew Valley Lake,
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
Сокол балобан на обучении (Falco cherrug).jpg, Falcon training in
Moscow Moscow ( , US chiefly ; rus, links=no, Москва, r=Moskva, p=mɐskˈva, a=Москва.ogg) is the capital and largest city of Russia. The city stands on the Moskva River in Central Russia, with a population estimated at 13.0 million ...
Falco cherrug Qatar.jpg, Captive bird,
Doha Doha ( ar, الدوحة, ad-Dawḥa or ''ad-Dōḥa'') is the capital city and main financial hub of Qatar. Located on the Persian Gulf coast in the east of the country, north of Al Wakrah and south of Al Khor, it is home to most of the coun ...
,
Qatar Qatar (, ; ar, قطر, Qaṭar ; local vernacular pronunciation: ), officially the State of Qatar,) is a country in Western Asia. It occupies the Qatar Peninsula on the northeastern coast of the Arabian Peninsula in the Middle East; it sh ...
Aa_saker_female.jpg, Captive bird Sakerfalke beim Wenden in der Luft - Zuschnitt.jpg, Flying bird at bird show


In culture

A Hungarian mythological bird, the
Turul The Turul is a mythological bird of prey, mostly depicted as a Falcon, in Hungarian tradition and Turkic tradition, and a national symbol of Hungarians. Origin The Turul is probably based on a large falcon. The Hungarian language word ''tur ...
, was probably a saker falcon (''kerecsensólyom''), and the saker falcon is the national bird of
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
. In 2012, the saker falcon was selected as the national bird of
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
. In
Disney The Walt Disney Company, commonly known as Disney (), is an American multinational mass media and entertainment conglomerate headquartered at the Walt Disney Studios complex in Burbank, California. Disney was originally founded on October ...
's ''
Mulan Hua Mulan () is a legendary folk heroine from the Northern and Southern dynasties era (4th to 6th century CE) of Chinese history. According to legend, Mulan took her aged father's place in the conscription for the army by disguising herself as ...
'', the Hun leader Shan Yu owns a saker falcon called Hayabusa. In the children's book ''Tiger Wars'' by
Steve Backshall Stephen James Backshall (born 21 April 1973) is an English naturalist, explorer, presenter and writer, best known for BBC TV's ''Deadly 60''. His other BBC work includes being part of the expedition teams in ''Lost Land of the Tiger'', ''Lo ...
, one of the main characters is called Saker after the falcon.


Use in falconry

The saker falcon has been used in falconry for thousands of years, and like its very close relative, the
gyrfalcon The gyrfalcon ( or ) (), the largest of the falcon species, is a bird of prey. The abbreviation gyr is also used. It breeds on Arctic coasts and tundra, and the islands of northern North America and the Eurosiberian region. It is mainly a reside ...
, is a highly regarded in it. Swift and powerful, it is effective against medium and large game bird species. Saker falcons can reach speeds of 120 to 150 km/h and suddenly swoop down on their prey. In recent years it is possible that hybrids of the saker falcon and peregrine falcon have been developed in order to provide falconers a bird with greater size and horizontal speed than the peregrine, with greater propensity for diving stoops on game than the saker.


References

* *


External links


saker.sciencefornature.org
*
Live 24hr view of a saker falcon nest
* * * *
Saker Falcon ''Falco cherrug'' Global Action Plan (SakerGAP)
{{Authority control
saker falcon The saker falcon (''Falco cherrug'') is a large species of falcon. This species breeds from central Europe eastwards across the Palearctic to Manchuria. It is mainly migratory except in the southernmost parts of its range, wintering in Ethiopia ...
Birds of Africa Birds of Central Asia Birds of Mongolia Birds of prey of Eurasia Birds of the Arabian Peninsula Birds of China Birds of Pakistan
saker falcon The saker falcon (''Falco cherrug'') is a large species of falcon. This species breeds from central Europe eastwards across the Palearctic to Manchuria. It is mainly migratory except in the southernmost parts of its range, wintering in Ethiopia ...
Falconry
saker falcon The saker falcon (''Falco cherrug'') is a large species of falcon. This species breeds from central Europe eastwards across the Palearctic to Manchuria. It is mainly migratory except in the southernmost parts of its range, wintering in Ethiopia ...