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''Sabal'' is a genus of palms (or fan-palms)
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the New World. Currently, there are 17 recognized species of ''Sabal'', including one hybrid species. The species are native to the subtropical and tropical regions of the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
, from the Gulf Coast/South Atlantic states in the
Southeastern United States The Southeastern United States, also referred to as the American Southeast or simply the Southeast, is a geographical region of the United States. It is located broadly on the eastern portion of the southern United States and the southern por ...
, south through the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
,
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
to
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
and
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. Members of this genus are typically identified by the leaves which originate from a bare, unarmed petiole in a fan-like structure. All members of this genus have a costa (or midrib) that extends into the leaf blade. This midrib can vary in length; and it is due to this variation that leaf blades of certain species of ''Sabal'' are strongly curved or strongly costapalmate (as in ''Sabal palmetto'' and ''Sabal etonia'') or weakly curved (almost flattened), weakly costapalmate, (as in ''Sabal minor''). Like many other palms, the fruit of ''Sabal'' are
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
, that typically change from green to black when mature.


Species

* ''Sabal'' × ''brazoriensis'' D.H.Goldman, Lockett & Read ''(S. minor × S. palmetto)'' -
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...


Prehistoric taxa

Extinct species within this genus include: * '' Sabal bigbendense'' Manchester et al. 2010† * ''
Sabal bracknellense ''Sabal'' is a genus of palms (or fan-palms) endemic to the New World. Currently, there are 17 recognized species of ''Sabal'', including one hybrid species. The species are native to the subtropical and tropical regions of the Americas, from th ...
'' (
Chandler Chandler or The Chandler may refer to: * Chandler (occupation), originally head of the medieval household office responsible for candles, now a person who makes or sells candles * Ship chandler, a dealer in supplies or equipment for ships Arts ...
) Mai
* '' Sabal grayana'' Brown 1962† * '' Sabal imperialis'' Brown 1962† * '' Sabal jenkinsii''† (
Reid Reid is a surname of Scottish origin. It means "red". People with the surname * Alan Reid (disambiguation) * Alex Reid (disambiguation), includes Alexander Reid * Amanda Reid, Australian Paralympic athlete * Amanda Reid (taxonomist), Australia ...
& Chandler)
Manchester Manchester () is a city in Greater Manchester, England. It had a population of 552,000 in 2021. It is bordered by the Cheshire Plain to the south, the Pennines to the north and east, and the neighbouring city of Salford to the west. The t ...


Formerly placed here

* ''
Serenoa repens ''Serenoa repens'', commonly known as saw palmetto, is the sole species currently classified in the genus ''Serenoa''. It is a small palm, growing to a maximum height around . It is endemic to the subtropical and tropical Southeastern United S ...
'' ( W.Bartram)
Small Small may refer to: Science and technology * SMALL, an ALGOL-like programming language * Small (anatomy), the lumbar region of the back * ''Small'' (journal), a nano-science publication * <small>, an HTML element that defines smaller text ...
(as ''S. serrulata'' (Michx.) Nutt. ex Schult. & Schult.f.)


Fossil record

These plants lived from the late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
to the
Quaternary The Quaternary ( ) is the current and most recent of the three periods of the Cenozoic Era in the geologic time scale of the International Commission on Stratigraphy (ICS). It follows the Neogene Period and spans from 2.58 million years ...
period (from 66 million to 12 thousand years ago). Fossils have been found in United States, Italy, Switzerland, Germany, Japan, the United Kingdom, and France.Paleobiology Database
/ref> Leaf fossils of †''Sabal lamanonis'' have been recovered from
rhyodacite Rhyodacite is a volcanic rock intermediate in composition between dacite and rhyolite. It is the extrusive equivalent of those plutonic rocks that are intermediate in composition between monzogranite and granodiorite. Rhyodacites form from rapid ...
tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock cont ...
of Lower
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
age in Southern
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
near the town of
Lučenec Lučenec (; german: Lizenz; hu, Losonc; yi, לאשאנץ; la, Lutetia HungarorumLelkes György (1992), Magyar helységnév-azonosító szótár, Balassi Kiadó, Budapest, 508 p.) is a town in the Banská Bystrica Region of south-central Slovaki ...
.27 million years old †''Sabal lamanonis'' and †''Sabal raphipholia'' leaf
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s in volcanic rocks have been described from the Evros region in
Western Thrace Western Thrace or West Thrace ( el, υτικήΘράκη, '' ytikíThráki'' ; tr, Batı Trakya; bg, Западна/Беломорска Тракия, ''Zapadna/Belomorska Trakiya''), also known as Greek Thrace, is a Geography, geograp ...
,
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
.


Phylogenetic work in ''Sabal'' (1990s – present)

The name ''Sabal'' was first applied to members of the group by
Michel Adanson Michel Adanson (7 April 17273 August 1806) was an 18th-century French botanist and naturalist who traveled to Senegal to study flora and fauna. He proposed a "natural system" of taxonomy distinct from the binomial system forwarded by Linnaeus. ...
in the 18th century. Previous names that this genus was associated with include ''
Corypha ''Corypha'' (gebang palm, buri palm or talipot palm) is a genus of palms (family Arecaceae), native to India, Malaysia, Indonesia, the Philippines, New Guinea and northeastern Australia (Cape York Peninsula, Queensland). They are fan palms (sub ...
'', ''
Chamaerops ''Chamaerops'' is a genus of flowering plants in the family Arecaceae. The only currently fully accepted species is ''Chamaerops humilis'', variously called European fan palm or the Mediterranean dwarf palm. It is one of the most cold-hardy p ...
'', ''
Rhapis ''Rhapis'' is a genus of about 10 species of small palms native to southeastern Asia from southern Japan and southern China south to Sumatra. The species are commonly known as lady palms. They are fan palms (subfamily Coryphoideae), with the l ...
''. This section highlights important
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
work done within the genus ''Sabal.'' In 1990, Scott Zona outlined key morphological and anatomical characters that he used to analyze species relationships of ''Sabal.'' Through this analysis of characters, Zona produced a
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
that portrays
evolutionary Evolution is change in the heredity, heritable Phenotypic trait, characteristics of biological populations over successive generations. These characteristics are the Gene expression, expressions of genes, which are passed on from parent to ...
relationships amongst 15 species of ''Sabal''. Based on the distribution of species within his cladogram, Zona recognized four distinct
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
s. The clades within his study include (Clade 1) ''
Sabal minor ''Sabal minor'', commonly known as the dwarf palmetto, is a small species of palm. It is native to the deep southeastern and south-central United States and northeastern Mexico. It is naturally found in a diversity of habitats, including maritime ...
; (''Clade 2) ''
Sabal bermudana ''Sabal bermudana'', commonly known as the Bermuda palmetto or bibby-tree, is one of 15 species of palm trees in the genus ''Sabal'' and is endemic to Bermuda although reportedly naturalized in the Leeward Islands. It was greatly affected by the ...
,
Sabal palmetto ''Sabal palmetto'' (, '' SAY-bəl''), also known as cabbage palm, cabbage palmetto, sabal palm, blue palmetto, Carolina palmetto, common palmetto, Garfield's tree, and swamp cabbage, is one of 15 species of palmetto palm. It is native to the So ...
,
Sabal miamiensis ''Sabal miamiensis,'' the Miami palmetto, is a rare plant species endemic to Dade County, Florida, in the vicinity of the city of Miami. Taxonomy The formal description of this as a new species was published in 1985, based largely on specim ...
,'' and ''
Sabal etonia ''Sabal etonia'', commonly known as the scrub palmetto is a species of palm. It is native only to peninsular Florida in the United States, where it is found in Florida sand pine scrub communities. Description ''Sabal etonia'' is a fan palm with ...
;'' (Clade 3) ''
Sabal maritima ''Sabal maritima'' is a species of palm which is native to Jamaica and Cuba. Description ''Sabal maritima'' is a fan palm with solitary, stout stems, which grows up to tall and in diameter. Plants have about 25 leaves, each with 70–110 leaf ...
,
Sabal domingensis ''Sabal domingensis'', the Hispaniola palmetto, is a species of palm which is native to Hispaniola (in both the Dominican Republic and Haiti) and Cuba. Description ''Sabal domingensis'' is a fan palm with solitary, very stout stems, which grow ...
,
Sabal causiarum ''Sabal causiarum'', commonly known as the Puerto Rico palmetto or Puerto Rican hat palm, is a species of Arecaceae, palm which is native to Hispaniola (in both the Dominican Republic and Haiti), Puerto Rico, and the British Virgin Islands. As it ...
, Sabal maurittiformis,
Sabal yapa ''Sabal yapa'' is a species of palm. It grows in Belize, Guatemala, western Cuba, and the Yucatan Peninsula region of Mexico (Yucatán, Campeche, Quintana Roo). It prefers limestone-based calcareous Calcareous () is an adjective meaning "mo ...
,
Sabal mexicana ''Sabal mexicana'' is a species of palm tree that is native to far southern North America. Common names include Rio Grande palmetto, Mexican palmetto, Texas palmetto, Texas sabal palm, palmetto cabbage and palma de mícharos. The specific epith ...
,'' and ''Sabal guatemalensis;'' (Clade 4) ''
Sabal uresana ''Sabal uresana'', commonly known as the Sonoran palmetto, is a species of Arecaceae, palm tree that is native to the foothills of the Sierra Madre Occidental in northwestern Mexico (states of Chihuahua (state), Chihuahua and Sonora). The specifi ...
, Sabal rosei, and Sabal pumos''. These clades associate closely with geographic distributions. All of the species within Clade 3 occur in the
Greater Antilles The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, and ...
and southern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, where species that occur in the
Greater Antilles The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, and ...
are more closely related to each other than those that occur in southern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. Although Clade 4 also occurs in
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, these species occur on the west coast where they are geographically separated from the
Mexican Mexican may refer to: Mexico and its culture *Being related to, from, or connected to the country of Mexico, in North America ** People *** Mexicans, inhabitants of the country Mexico and their descendants *** Mexica, ancient indigenous people ...
species within the southern part of the country. The remaining two clades, Clade 1 and Clade 2 predominantly occur in the
southeastern United States The Southeastern United States, also referred to as the American Southeast or simply the Southeast, is a geographical region of the United States. It is located broadly on the eastern portion of the southern United States and the southern por ...
although ''S. palmetto'' and ''S. minor'' are also known from
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
and the
Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the Atlantic Ocean, North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to ...
''(S. palmetto)'' and northern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
''(S. minor).'' ''Sabal bermudana'' is only known from the
Bermuda Islands ) , anthem = "God Save the King" , song_type = National song , song = "Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , es ...
. In 2016 Heyduk, Trapnell, Barrett, and Leebens-Mack conducted a new study on ''Sabal'' that analyzed molecular (e.g. nuclear, plastid) data from 15 species of the group. This study incorporated
plastid The plastid (Greek: πλαστός; plastós: formed, molded – plural plastids) is a membrane-bound organelle found in the Cell (biology), cells of plants, algae, and some other eukaryotic organisms. They are considered to be intracellular endosy ...
and nuclear sequence data that together were used to estimate the relatedness between the species of ''Sabal.'' The results of the study show species relationships to be different from the distribution of Zona's cladogram. Within the framework of this study, a major difference between the results of Zona and this study is the placement of "Clade 4" (''
Sabal uresana ''Sabal uresana'', commonly known as the Sonoran palmetto, is a species of Arecaceae, palm tree that is native to the foothills of the Sierra Madre Occidental in northwestern Mexico (states of Chihuahua (state), Chihuahua and Sonora). The specifi ...
, Sabal rosei, and Sabal pumos'') which split and integrate these species throughout the
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
of ''Sabal.'' The largest of the clades identified by Zona, "Clade 3" is disrupted significantly as it is split into multiple clades. Although ''Sabal causiarum'' and ''S. domingensis'' retain their relationship as
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
, they are included in a clade that also includes ''S. maritima'' and ''S. rosei.'' Despite these disruptions in placement between these two studies, the overall integrity of "Clade 1" and "Clade 2" is in congruence with the clades established from the molecular data.


Pollinators and parasitoids

''Sabal'' species are used as food sources by several species of birds including '' Mimus polyglottos,
Turdus migratorius The American robin (''Turdus migratorius'') is a migratory bird of the true thrush genus and Turdidae, the wider thrush family. It is named after the European robin because of its reddish-orange breast, though the two species are not closely ...
, Dendroica coronata,
Corvus ossifragus The fish crow (''Corvus ossifragus'') is a species of crow associated with wetland habitats in the eastern and southeastern United States. Taxonomy and etymology The fish crow was given its binomial name by the Scottish ornithologist Alexand ...
,'' and '' Drycopus pileatus'', '' Caryobruchus'', and various species of
hymenoptera Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...
. Bears (''
Ursus americanus Ursus is Latin for bear. It may also refer to: Animals *Ursus (mammal), ''Ursus'' (mammal), a genus of bears People * Ursus of Aosta, 6th-century evangelist * Ursus of Auxerre, 6th-century bishop * Ursus of Solothurn, 3rd-century martyr * Ursus ...
)'' and racoons are also known to feed on fruit of various species of ''Sabal. Sabal palmetto'' is recorded to have its own lichen, '' Arthonia rubrocincta,'' that only occurs on the leaf bases of the Cabbage Palm (''Sabal palmetto''). In Europe, the introduced Lepidopteran species ''
Paysandisia archon ''Paysandisia archon'' is a moth of the family Castniidae. It is native to Uruguay and central Argentina and has been accidentally introduced to Europe, where it is spreading rapidly. It is considered the only member of the genus ''Paysandisia''. ...
'' has become a prominent pest whose
larvae A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
are known to feed on some of the cultivated species of ''Sabal.''


Uses

Arborescent species are ofte
transplanted
from natural stands into urban landscapes and are rarely grown in nurseries due to slow growth. Several species are cultivated as
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...
s and because several species are relatively cold-hardy, can be grown farther north than most other palms. The central bud of ''Sabal'' ''palmetto'' is edible and, when cooked, is known a
'swamp cabbage'
Mature fronds are used as thatch, to make straw hats, and for weaving mats.


Symbolic use

A
silhouette A silhouette ( , ) is the image of a person, animal, object or scene represented as a solid shape of a single colour, usually black, with its edges matching the outline of the subject. The interior of a silhouette is featureless, and the silhou ...
of a palmetto (''S. palmetto'') appears on the official
flag of South Carolina The flag of South Carolina is a symbol of the U.S. state of South Carolina consisting of a blue field with a white palmetto tree and white crescent. Roots of this design have existed in some form since 1775, being based on one of the first Re ...
.Netstate, South Carolina State Flag
/ref> Two images of ''S. palmetto'' appear on the
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
br>state seal
''Sabal palmetto'' is the state tree of bot
Florida
and South Carolina.


References


External links


''Sabal'' images
at Fairchild Tropical Botanical Gardens

at Scanpalm {{Taxonbar, from1=Q132826, from2=Q21447238 Arecaceae genera Neotropical realm flora Taxa named by Michel Adanson