Rubus

''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, with over 1,350 species.

Raspberries, blackberries, and dewberries are common, widely distributed members of the genus. Most of these plants have woody stems with prickles like roses; spines, bristles, and gland-tipped hairs are also common in the genus. The ''Rubus'' fruit, sometimes called a bramble fruit, is an aggregate of drupelets. The term "cane fruit" or "cane berry" applies to any ''Rubus'' species or hybrid which is commonly grown with supports such as wires or canes, including raspberries, blackberries, and hybrids such as loganberry, boysenberry, marionberry and tayberry. The stems of such plants are also referred to as canes.

Description

Most species in the genus are hermaphrodites, '' Rubus chamaemorus'' being an exception.

''Rubus'' species have a basic chromosome number of seven. Polyploidy from the diploid (14 chromosomes) to the tetradecaploid (98 chromosomes) is exhibited.

Taxonomy

Modern classification

''Rubus'' is very complex, particularly within the blackberry/dewberry subgenus (''Rubus''), with polyploidy, hybridization, and facultative apomixis apparently all frequently occurring, making species classification of the great variation in the subgenus one of the grand challenges of systematic botany.

Some treatments have recognized dozens of species each for what other, comparably qualified botanists have considered single, more variable species. On the other hand, species in the other ''Rubus'' subgenera (such as the raspberries) are generally distinct, or else involved in more routine one-or-a-few taxonomic debates, such as whether the European and American red raspberries are better treated as one species or two (in this case, the two-species view is followed here, with '' R. idaeus'' and '' R. strigosus'' both recognized; if these species are combined, then the older name ''R. idaeus'' has priority for the broader species).

The classification presented below recognizes 13 subgenera within ''Rubus'', with the largest subgenus (''Rubus'') in turn divided into 12 sections. Representative examples are presented, but many more species are not mentioned here. A comprehensive 2019 study found subgenera ''Orobatus'' and ''Anoplobatus'' to be monophyletic, while all other subgenera to be paraphyletic or polyphyletic.

Phylogeny

The genus has a likely North American origin, with fossils known from the Eocene-aged Florissant Formation of Colorado. ''Rubus'' expanded into Eurasia, South America, and Oceania during the Miocene. Fossil seeds from the early Miocene of ''Rubus'' have been found in the Czech part of the Zittau Basin. Many fossil fruits of †''Rubus laticostatus'', †''Rubus microspermus'' and †''Rubus semirotundatus'' have been extracted from bore hole samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin, West Carpathians, Poland.

Molecular data have backed up classifications based on geography and chromosome number, but following morphological data, such as the structure of the leaves and stems, do not appear to produce a phylogenetic classification.

Species

Better-known species of ''Rubus'' include:

*'' Rubus ablatus''
*'' Rubus aboriginum'' – garden dewberry
*''Rubus allegheniensis'' – Allegheny blackberry
*'' Rubus arcticus'' – Arctic raspberry
*''Rubus argutus''
*''Rubus armeniacus'' – Himalayan blackberry
*'' Rubus caesius'' – European dewberry
*'' Rubus canadensis'' – Canadian blackberry
*'' Rubus chamaemorus'' – cloudberry
*''Rubus cockburnianus''
*''Rubus coreanus'' – bokbunja
*''Rubus crataegifolius''
*''Rubus deliciosus''
*''Rubus domingensis''
*''Rubus ellipticus''
*'' Rubus flagellaris'' – northern dewberry
*'' Rubus fraxinifolius'' – mountain raspberry
*'' Rubus glaucus''
*''Rubus hawaiensis''
*'' Rubus hayata-koidzumii''
*''Rubus hispidus'' – swamp dewberry
*''Rubus idaeus'' – European red raspberry
*''Rubus illecebrosus''
*''Rubus laciniatus'' – cutleaf evergreen blackberry
*''Rubus leucodermis'' – whitebark raspberry or western black raspberry
*'' Rubus moluccanus''
*'' Rubus nepalensis''
*'' Rubus nivalis''
*''Rubus niveus''
*'' Rubus occidentalis'' – black raspberry
*'' Rubus odoratus'' – flowering raspberry
*''Rubus parviflorus'' – thimbleberry
*'' Rubus parvifolius'' – small-leaf bramble (Australia)
*'' Rubus pedatus''
*'' Rubus pensilvanicus'' – Pennsylvania blackberry
*'' Rubus phoenicolasius'' – wine raspberry or wineberry
*''Rubus plicatus''
*'' Rubus probus''
*'' Rubus pubescens'' – dwarf red blackberry
*'' Rubus rosifolius''
*'' Rubus saxatilis'' – stone bramble
*''Rubus spectabilis'' – salmonberry
*''Rubus strigosus'' – American red raspberry
*'' Rubus tricolor''
*''Rubus ulmifolius''
*''Rubus ursinus'' – trailing blackberry

A more complete subdivision is as follows:

Hybrid berries

The term "hybrid berry" is often used collectively for those fruits in the genus ''Rubus'' which have been developed mainly in the U.S. and U.K. in the last 130 years. As ''Rubus'' species readily interbreed and are apomicts (able to set seed without fertilisation), the parentage of these plants is often highly complex, but is generally agreed to include cultivars of blackberries (''R. ursinus'', ''R. fruticosus'') and raspberries (''R. idaeus''). The British National Collection of ''Rubus'' stands at over 200 species and, although not within the scope of the National Collection, also hold many cultivars.

The hybrid berries include:-
* loganberry (California, U.S., 1883) ''R.'' × ''loganobaccus'', a spontaneous hybrid between ''R. ursinus'' 'Aughinbaugh' and ''R. idaeus'' 'Red Antwerp'
* boysenberry (U.S., 1920s) a hybrid between ''R. idaeus'' and ''R.'' × ''loganobaccus''
* olallieberry (U.S., 1930s) a hybrid between the loganberry and youngberry, themselves both hybrid berries
* veitchberry (Europe, 1930s) a hybrid between ''R. fruticosus'' and ''R. idaeus''
* skellyberry (Texas, U.S., 2000s), a hybrid between ''R. invisus'' and ''R. phoenicolasius''
* marionberry (1956) now thought to be a blackberry cultivar ''R.'' 'Marion'
* silvanberry, ''R.'' 'Silvan', a hybrid between ''R.'' 'Marion' and the boysenberry
* tayberry (Dundee, Scotland, 1979), another blackberry/raspberry hybrid
* tummelberry, ''R.'' 'Tummel', from the same Scottish breeding programme as the tayberry
* hildaberry (1980s), a tayberry/boysenberry hybrid discovered by an amateur grower
* youngberry, a complex hybrid of raspberries, blackberries, and dewberries

Etymology

The generic name means blackberry in Latin and was derived from the word ''ruber'', meaning "red".

The blackberries, as well as various other ''Rubus'' species with mounding or rambling growth habits, are often called brambles. However, this name is not used for those like the raspberry that grow as upright canes, or for trailing or prostrate species, such as most dewberries, or various low-growing boreal, arctic, or alpine species. The scientific study of brambles is known as "batology".

See also

* List of Lepidoptera that feed on ''Rubus''
* Mulberry, an unrelated deciduous tree with similar looking fruit

References

External links

*

''Rubus''
at the Western Kentucky University

{{Authority control

Rosaceae genera
Subshrubs
Extant Eocene first appearances